Akihiko Shinohara

Review of the sawfly genus Empria (Hymenoptera, Tenthredinidae) in Japan

Abstract The following eleven Empria species are reported from Japan: Empria candidata (Fallén, 1808), Empria japonica Heidemaa & Prous, 2011, Empria liturata (Gmelin, 1790), Empria loktini Ermolenko, 1971, Empria plana (Jakowlew, 1891), Empria quadrimaculata Takeuchi, 1952, Empria rubicola Ermolenko, 1971, Empria tridens (Konow, 1896), Empria tridentis Lee & Ryu, 1996, Empria honshuana Prous & Heidemaa, sp. n., and Empria takeuchii Prous & Heidemaa, sp. n. The lectotypes of Poecilosoma pallipes Matsumura, 1912, Empria itelmena Malaise, 1931, Tenthredo candidata Fallén, 1808, and Tenthredo (Poecilostoma) hybrida Erichson, 1851 are designated. Empria itelmena Malaise, 1931, syn. n. is synonymized with Empria plana (Jakowlew, 1891). Poecilosoma pallipes Matsumura, 1912, previously assigned to Empria, is transferred to Monsoma, creating Monsoma pallipes (Matsumura, 1912), comb. n. Results of phylogenetic analyses using mitochondrial (COI) and nuclear (ITS1 and ITS2) sequences are also provided.

Xiphydriid Woodwasps (Hymenoptera: Xiphydriidae) of Korea

Abstract. Six species in three genera of Xiphydriidae are recorded from Korea, Euxiphydria potanini (Jakovlev, 1891), Xiphydriola amurensis Semenov, 1921, Xiphydria annulitibia Takeuchi, 1936, X. camelus (Linnaeus, 1758), X. jakovlevi Semenov and Gussakovskij, 1935, and X. palaeanarctica Semenov, 1921. Xiphydriola amurensis, Xiphydria annulitibia, and X. jakovlevi are new country records. The following taxonomic changes are proposed: Xiphydriola nishijimai (Togashi, 1998), n. comb., and Nasoxiphia Maa, 1949 = Xiphydria Latreille, 1803, n. syn.

The East Asian Xyela species (Hymenoptera: Xyelidae) associated with Japanese Red Pine ( Pinus densiflora ; Pinaceae) and their distribution history

The association of five East Asian species of Xyela Dalman, 1819 with Pinus densiflora Siebold & Zuccarini, 1842 has been demonstrated by the identification of reared adults or of adults collected by emergence traps. The host plant of X. japonica Rohwer, 1910 is doubtful. These Xyela species occur as sibling species or at least as pairs of morphologically similar species, which are distributed vicariantly on the East Asian mainland and in Japan, respectively. The vicariance events for the Korean-Japanese species pairs date back to the disconnection of ancient Japan and South-East Asia during the Mindel-Riss interglacial period (320,000-180,000 BP). The most southern record of a xyelid sawfly from South-East Asia is reported for an unidentified Xyela female from southern Vietnam. Xyela densiflorae Blank & Shinohara (South Korea), X. occidentalis Blank & Shinohara (northeastern China, South Korea), Xyela par Blank & Shinohara (South Korea), and X. tecta Blank & Shinohara (Japan) are described. X. variegata Rohwer, 1910, formerly regarded as a synonym of X. julii (Brebisson, 1818), is reestablished. The male of X. japonica is described for the first time. A key for the identification of these species is presented.

Pachyprotasis kojimai sp. nov.—the “ Pachyprotasis nigronotata ” of Japanese authors (Hymenoptera: Tenthredinidae)

The species formerly known as Pachyprotasis nigronotata Kriechbaumer, 1874 in Japan is described as P. kojimai Taeger & Shinohara, sp. nov. Both taxa are not very closely related, but were hitherto mixed up because of their similar coloration. The new species is endemic to the mountains of central Honshu and P. nigronotata is to be deleted from the fauna of Japan.

The webspinning sawfly genus Neurotoma (Hymenoptera, Pamphiliidae) of South Korea: Neurotoma silla sp. nov. and a key to species

Neurotoma silla sp. nov. is described and illustrated from South Korea. It belongs to the N. saltuum subgroup of the N. nemoralis group and is well characterized by the large size, the mostly black body with mostly black antennae and partly pale yellow legs, the densely punctate but nearly glabrous head with smooth interspaces between punctures, and the sharply carinate occipital carina. Six species of Neurotoma are known now from South Korea, including three species endemic to the country. A key to the South Korean species of Neurotoma is given.

DNA Barcodes Identify the Larvae and Unassociated Male of Three Onycholyda Sawflies (Hymenoptera, Pamphiliidae) from China

A molecular analysis based on mitochondrial cytochrome oxidase subunit 1 (COI) gene sequences has indicated that larvae collected in Sichuan and Zhejiang Provinces, China, belong to Onycholyda xanthogaster Shinohara, 1999, and O. fulvicornis Shinohara, in Shinohara Wei, 2016 (Hymenoptera, Pamphiliidae), and that a male Onycholyda specimen from Mt. Tianmushan, Zhejiang Province is the hitherto unknown male of O. tianmushana Shinohara Xiao, 2006. The first host plant records are Rubus inopertus (Focke) Focke (Rosaceae) for O. xanthogaster and Rubus hirsutus Thunb. for O. fulvicornis. The larvae of O. xanthogaster and O. fulvicornis are briefly described and O. xanthogaster is newly recorded from Sichuan Province. The male of O. tianmushana is described for the first time.

Figures 7–13 In The Webspinning Sawfly Genus Neurotoma (Hymenoptera, Pamphiliidae) Of South Korea: Neurotoma Silla Sp. Nov. And A Key To Species

FIGURES 7–13. Neurotoma silla (paratype, male). 7, Habitus in dorsal view; 8, habitus in ventral view; 9, head in frontal view; 10, apex of abdomen in dorsal view; 11, habitus in lateral view; 12, apex of abdomen in ventral view; 13, antenna. Scale bars: 7, 8, 11, 2 mm; 9, 13, 1 mm; 10, 12, 0.5 mm.

Figure 5 In Pachyprotasis Kojimai Sp. Nov. — The " Pachyprotasis Nigronotata " Of Japanese Authors (Hymenoptera: Tenthredinidae)

FIGURE 5. The maximum likelihood tree (T 92 + G) showing relationships among 17 samples of 11 species of Pachyprotasis. Numbers on branches are bootstrap values (1000 bootstrap replicates), only values> 50 % are shown.

Revision of Neocolochelyna Malaise (Hymenoptera: Tenthredinidae).

Neocolochelyna Malaise, 1937 is revised. Curvatapenis Wei, 2002 is treated as a subgenus of Neocolochelyna. The genus and its type species, N. montana Konow, 1898, as well as N. (C.) itoi Takeuchi, 1951 and N. (C.) testaceoa Wei, 2002 are redescribed based on types and new material. N. (N.) acutilabria Niu & Wei sp. nov., N. (C.) hyalina Niu & Wei sp. nov. and N. (C.) wui Niu & Wei sp. nov. are described. Neocolochelyna hakusana Togashi, 1995 is a new synonym (syn. nov.) of N. itoi Takeuchi, 1951. Neocolochelyna (Curvatapenis) testaceoa (Wei, 2002) is a new combination (comb. nov.). A key to species of Neocolochelyna is provided.

Leaf-rolling Sawflies of the Pamphilius vafer complex (Hymenoptera, Pamphiliidae) in Europe

Shinohara, A.: Leaf-rolling Sawflies of the Pamphilius vafer complex (Hymenoptera, Pamphiliidae) in Europe. Insect Syst. Evol. 34: 453-480. Copenhagen, December 2003. ISSN 1399-560X.Pamphiliusvafer complex of species includes four species in Europe; P. vafer (Linnaeus, 1767) [=Tenthredo depressa Schrank, 1781] from Europe (including Britain) and Siberia?, P. pallipes (Zetterstedt, 1838) [=Lyda flavipes Zetterstedt, 1838, =Lyda variegata Zaddach, 1866] from Europe and Siberia?, P. nigrifemoratus Shinohara & Taeger, 1990, from the eastern Carpathians, and P. kontuniemii sp. n. [=P. depressus of Kontuniemi, 1949, and Viitasaari, 1982, not Schrank, 1781] from Europe (not including Britain). The last mentioned new species, closely resembling P. vafer, is described and illustrated. A neotype is designated for Tenthredo depressa Schrank, 1781, and lectotypes are designated for Lyda pallipes Zetterstedt, 1838, and Lyda flavipes Zetterstedt, 1838. Each species is diagnosed and a key is given to separate the four species. Previous distribution and host plant records are critically reviewed for each species.