Christopher G. Maples

Ichnology of an Upper Carboniferous fluvio-estuarine paleovalley: The Tonganoxie Sandstone, Buildex Quarry, Eastern Kansas, USA

Tidal rhythmites of the Tonganoxie Sandstone Member (Stranger Formation, Douglas Group) at Buildex Quarry, eastern Kansas, contain a relatively diverse ichnofauna. The assemblage includes arthropod locomotion (Dendroidichnites irregulare, Diplichnites gouldi types A and B, Diplopodichnus biformis, Kouphichnium isp., Mirandaichnium famatinense, and Stiaria intermedia), resting (Tonganoxichnus buildexensis) and feeding traces (Stiallia pilosa, Tonganoxichnus ottawensis); grazing traces (Gordia indianaensis, Helminthoidichnites tenuis, Helminthopsis hieroglyphica); feeding structures (Circulichnis montanus, Treptichnus bifurcus, Treptichnus pollardi, irregular networks), fish traces (Undichna britannica, Undichna simplicitas), tetrapod trackways, and root traces. The taxonomy of some of these ichnotaxa is briefly reviewed and emended diagnoses for Gordia indianaensis and Helminthoidichnites tenuis are proposed. Additionally, the combined name Dendroidichnites irregulare is proposed for nested chevron trackways. Traces previously regarded as produced by isopods are reinterpreted as myriapod trackways (D. gouldi type B). Trackways formerly interpreted as limulid crawling and swimming traces are assigned herein to Kouphichnium isp and Dendroidichnites irregulare, respectively. Taphonomic analysis suggests that most grazing and feeding traces were formed before the arthropod trackways and resting traces. Grazing/feeding traces were formed in a soft, probably submerged substrate. Conversely, the majority of trackways and resting traces probably were produced subaerially in a firmer, dewatered and desiccated sediment. The Buildex Quarry ichnofauna records the activity of a terrestrial and freshwater biota. The presence of this assemblage in tidal rhythmites is consistent with deposition on tidal flats in the most proximal zone of the inner estuary, between the maximum landward limit of tidal currents and the salinity limit further towards the sea.

Contrasting depositional models for Pennsylvanian black shale discerned from molybdenum abundances

Some black shales contain >1,000 ppm Mo, comparable to conventional ores. These include Mo-rich Pennsylvanian black shales that were deposited rapidly near shore as epeiric seas inundated peat swamps, while coeval beds, less enriched in Mo, accumulated slowly offshore. Concentration of Mo probably resulted from retention of the metal by organic matter in acidic pore fluids characteristic of nearshore deposits. Thus, in Pennsylvanian shales where organic productivity is significant, Mo variations may be used to distinguish nearshore from offshore conditions. Enrichments in Mo occur in black shales of other ages; therefore a high Mo content may be useful as a general criterion for nearshore environments and other conditions with strong pH gradients.

Taxonomic reassessment of the ichnogenus beaconichnus and additional examples from the Carboniferous of Kansas, U.S.A.

The ichnogenus Beaconichnus (Gevers 1973), an arthropod trace fossil, includes very different forms that comprise five ichnospecies, namely B. darwi‐nunt (Gevers 1971), B. gouldi (Gevers 1971), B. ahtarcticum (Gevers 1971), B. giganteum Gevers and Twomey 1982, and B. wrrighti Gevers and Twomey 1982. The original diagnosis of Beaconichnus is rather vague and potentially may accomodate virtually every arthropod trackway described from the fossil record. In view of these problems, the validity of Beaconichnus is reassessed and each of its ichnospecies is reviewed. We conclude that B. darwinum is a junior synonym of Diplopodichnus biformis Brady 1947; B. antarcticum should be regarded as Palmich‐niunt antarcticum; and B. wrighti is a nomen nudum. Additionally, we agree with previous proposals in considering B. gouldi as the senior synonym of B. giganteum, and including it in Diplichnites Dawson 1873. Therefore, we suggest that the ichnogenus Beaconichnus is best disregarded. Additionally, we describe specimen...

The ichnogenus Curvolithus revisited

The ichnogenus Curvolithus Fritsch, 1908, originally described from the Ordovician of the Prague Basin, typically comprises ribbonlike or tonguelike, flattened, endostratal traces with three rounded lobes on the upper surface. However, considerable confusion persists regarding the ichnotaxonomic status and diagnostic features of its ichnospecies. The type specimens of this ichnotaxon, overlooked in most subsequent reports, are redescribed herein. Curvolithus multiplex Fritsch, 1908, the type species, is retained for specimens with a trilobate upper surface and a quadralobate lower surface, in contrast to the criteria adopted by subsequent authors. The other ichnospecies originally proposed from the type locality, C. gregarius Fritsch, 1908, actually consists of a series of grouped parallel scratch marks forming ridges and should be removed from Curvolithus. Subsequently, four ichnospecies were defined: C.? davidis Webby 1970; C. annulatus Badve and Ghare 1978; C. aequus Walter et al. 1989; and C. manitouensis Maples and Suttner 1990. Curvolithus? davidis shows the typical trilobation of Curvolithus apparently in its lower surface, but the morphology of the upper surface is uncertain. Accordingly, it does not warrant ichnospecific assessment, and is regarded as a nomen dubium. The nature of the annulations on the trilobate upper surface of C. annulatus is unclear, and this ichnospecies is also best considered as a nomen dubium. Curvolithus aequus has a bilobate lower surface and probably represents washed out specimens of Didymaulichnus. Finally, C. manitouensis comprises specimens with a smooth, trilobate upper surface and a smooth, quadralobate lower surface, and is best regarded as a junior synonym of C. multiplex. Curvolithus multiplex has been used incorrectly for Curvolithus with a trilobate upper surface and a trilobate to unilobate lower surface. The new ichnospecies, Curvolithus simplex, is proposed herein for such traces. Curvolithus is interpreted as a locomotion trace (Repichnia) of endostratal carnivores, possibly gastropods, flatworms, or nemerteans. Curvolithus is a component of the Cruziana ichnofacies in shallow-marine facies, either of normal salinity or slightly brackish, in the latter case typically associated with fan deltas.

The Hamilton Konservat-Lagerstätte: Stephanian terrestrial biota in a marginal-marine setting

The Stephanian Konservat-Lagerstatte of Hamilton, Kansas, deposited in a marine-estuarine environment preserves a mixture of terrestrial, fresh-water, and marine fossil organisms. The marine component is the most diverse taxonomically, whereas one ostracod species, commonly interpreted as a fresh-water form, together with vascular plant debris dominates volumetrically. Well-preserved terrestrial and aquatic vertebrates and arthropods were embedded in rapidly deposited calcareous mud conducive to microbial early-diagenesis of vertebrate soft tissues in a tidal estuarine setting. Many vertebrate fossils show no evidence of preburial decay or disarticulation

Origin of the Bear Gulch Beds (Namurian, Montana, USA)

Abstract The Bear Gulch Beds of the Heath Formation are well known for their diverse and well-preserved assemblage offish, arthropods, and soft-bodied animals (they constitute a Lagerstatte). The Bear Gulch is a lens of lithographic limestone (approximately 12 km in lateral extent and up to 30 m thick) surrounded by black, platy shale. The lens is composed primarily of alternating massive beds (up to 25 cm thick) and argillaceous platy beds (up to 30 cm thick). Platy and massive beds are both composed primarily of normally graded laminations (1–15 mm thick). Laminations typically have sharp bases and grade upward from microspar to micrite. Lateral continuity of individual beds (at least 1 km) and laminations (at least 500 m), lack of evidence of bottom currents, and paucity of erosional features all suggest a quiet-water environment. Fossils are generally rare in the Bear Gulch Beds. The most common fossils in most beds are cephalopods, shrimp, fish, and soft-bodied organisms. Rare beds contain abundant benthic fossils including brachiopods, sponges, bivalves, conulariids, and crinoids. Fish preservation ranges from completely articulated with traces of internal organs and preserved color patterns to completely disarticulated, however most fish are well preserved. Fish and shrimp occur at the boundaries between laminations. Preservational quality of fossils and presence of abundant dewatering structures suggest rapid deposition. Lack of normal-marine shelly fossils in most beds may indicate stressful conditions at the seafloor, however some fish and shrimp were apparently well adapted for a benthic habitat. No evidence of current-modified fish debris has been observed and only rarely are fish aligned on a single bedding plane. Conditions at the seafloor were calm, possibly inhospitable, but frequently disturbed by rapid depositional events. The mechanism of deposition of the laminations remains enigmatic.

Response of selected binomial coefficients to varying degrees of matrix sparseness and to matrices with known data interrelationships

Numerous departures from ideal relationships are revealed by Monte Carlo simulations of widely accepted binomial coefficients. For example, simulations incorporating varying levels of matrix sparseness (presence of zeros indicating lack of data) and computation of expected values reveal that not only are all common coefficients influenced by zero data, but also that some coefficients do not discriminate between sparse or dense matrices (few zero data). Such coefficients computationally merge mutually shared and mutually absent information and do not exploit all the information incorporated within the standard 2 × 2 contingency table; therefore, the commonly used formulae for such coefficients are more complicated than the actual range of values produced. Other coefficients do differentiate between mutual presences and absences; however, a number of these coefficients do not demonstrate a linear relationship to matrix sparseness. Finally, simulations using nonrandom matrices with known degrees of row-by-row similarities signify that several coefficients either do not display a reasonable range of values or are nonlinear with respect to known relationships within the data. Analyses with nonrandom matrices yield clues as to the utility of certain coefficients for specific applications. For example, coefficients such as Jaccard, Dice, and Baroni-Urbani and Buser are useful if correction of sparseness is desired, whereas the Russell-Rao coefficient is useful when sparseness correction is not desired.