David Rubilar-Rogers

Aristonectes quiriquinensis, sp. nov., A New Highly Derived Elasmosaurid from the Upper Maastrichtian of Central Chile

ABSTRACT This paper describes a new species of elasmosaurid plesiosaur, Aristonectes quiriquinensis, sp. nov., based on a partial skeleton recovered from upper Maastrichtian beds of the Quiriquina Formation of central Chile. The material described here consists of two skeletons, one collected near the village of Cocholgue, and a second juvenile specimen from Quiriquina Island. Prior to these finds, Aristonectes was viewed as a monospecific genus, including only the enigmatic Aristonectes parvidens, the holotype of which consists of an incomplete skull and incomplete postcranium. Other material referred to the genus includes an incomplete juvenile skull and other postcranial material from the upper Maastrichtian of Antarctica, as well as a partial skull from the Quiriquina Formation of central Chile. The relationships of Aristonectes have been controversial, with competing theories assigning the genus to Cryptoclididae, Elasmosauridae, and Aristonectidae; however, there is a developing consensus that Aristonectes is a derived elasmosaurid, and this paper gives strong evidence for this view. Comparison of the specimen here studied with the holotype of A. parvidens demonstrates that A. quiriquinensis is a distinct species. The completeness of the adult skeleton allows the first confident size estimates for adult Aristonectes. It is a large plesiosaurian with a relatively large skull with numerous homodont teeth, a moderately long and laterally compressed neck, and relatively narrow trunk, with slender and elongate forelimbs. The two specimens are restricted to the upper Maastrichtian of central Chile, posing questions concerning the austral circumpolar distribution of different elasmosaurids towards the end of the Cretaceous.

An enigmatic plant-eating theropod from the Late Jurassic period of Chile

Theropod dinosaurs were the dominant predators in most Mesozoic era terrestrial ecosystems. Early theropod evolution is currently interpreted as the diversification of various carnivorous and cursorial taxa, whereas the acquisition of herbivorism, together with the secondary loss of cursorial adaptations, occurred much later among advanced coelurosaurian theropods. A new, bizarre herbivorous basal tetanuran from the Upper Jurassic of Chile challenges this conception. The new dinosaur was discovered at Aysén, a fossil locality in the Upper Jurassic Toqui Formation of southern Chile (General Carrera Lake). The site yielded abundant and exquisitely preserved three-dimensional skeletons of small archosaurs. Several articulated individuals of Chilesaurus at different ontogenetic stages have been collected, as well as less abundant basal crocodyliforms, and fragmentary remains of sauropod dinosaurs (diplodocids and titanosaurians).

Theropod Dinosaur Trackways from the Lower Cretaceous of the Chacarilla Formation, Chile

We describe sixteen theropod dinosaur trackways from site III (Lower Cretaceous?) of the Quebrada Cha- carilla in the Chacarilla Formation (Jurassic-Lower Cretaceous), northern Chile. The trackbed belongs to a meandering river environment, with recurrent track assemblages. We fi nd that the main direction of the ichnites from site III parallels that of water fl ow. Two theropod footprint morphotypes are recognized: A) digit II and IV occupying about 0.5 times the total footprint length, and B) digit II and IV occupying less than 0.35 times the total footprint length. We estimate similar speed (~7 km/h) for the two parallel theropod trackways of morphotype A, suggesting synchro nized walking. We also describe some of the largest theropod footprints from South America (>60 cm), which are all of morphotype B.

A new species of chimaeriform (Chondrichthyes, Holocephali) from the uppermost Cretaceous of the Lopez de Bertodano Formation, Isla Marambio (Seymour Island), Antarctica

Abstract We describe a new chimaeriform fish, Callorhinchus torresi sp. nov., from the uppermost Cretaceous (late Maastrichtian) of the López de Bertodano Formation, Isla Marambio (Seymour Island), Antarctica. The material shows it is distinct from currently known fossil and extant species of the genus, whereas the outline of the tritors (abrasive surfaces of each dental plate) shows an intermediate morphology between earlier records from the Cenomanian of New Zealand and those from the Eocene of Isla Marambio. This suggests an evolutionary trend in tritor morphology in the lineage leading to modern callorhynchids, during the Late Cretaceous-Palaeogene interval.

A new titanosaur sauropod from the Atacama Desert, Chile

Partial remains of a titanosaur sauropod collected in the Tolar Formation (Upper Cretaceous) at the Atacama Desert (Antofagasta Region), northern Chile, is described, and a new species, Atacamatitan chilensis gen. et sp. nov., is erected. The material consists mainly of dorsal and caudal vertebrae, part of a humerus and a femur. The presence of a titanosaur confirms the Cretaceous age for the outcrops of red sandstone of the Tolar Formation whose age was previously uncertain, ranging from the Upper Cretaceous to the Paleocene. The new specimen represents the most complete dinosaur reported for this region and one of the most complete titanosaur known from Chile and the pacific margin of South America so far.

NEW CHONDRICHTHYANS FROM THE UPPER CRETACEOUS (CAMPANIAN-MAASTRICHTIAN) OF SEYMOUR AND JAMES ROSS ISLANDS, ANTARCTICA

Abstract We present new records of chondrichthyans recovered from strata of Maastrichtian age of the López de Bertodano Formation, Seymour (=Marambio) Island, and from levels of latest Campanian age of the Santa Marta Formation, James Ross Island, both located in the eastern Antarctic Peninsula. The material from Marambio Island comprises an associated assemblage with the first records of an indeterminate odontaspidid different from Odontaspis, as well as the genera Pristiophorus, Squatina, Paraorthacodus, and the species Chlamydoselachus tatere from the López de Bertodano Formation. Also, the studied section provides a well-constrained age for several taxa already recognized in the López de Bertodano Formation only by scattered samples of Maastrichtian age for the first time. The assemblage from Marambio Island is representative of one of the latest environmental conditions during the end of the Cretaceous in the coastal seas of the Larsen Basin before major changes that began after the K/P boundary. In addition, the finds from James Ross Island comprise the southernmost records of the neoselachians Cretalamna sp., Centrophoroides sp., as well as the holocephalans Callorhinchus sp. and an indeterminate rhinochimaerid, extending the occurrence of some of these taxa into the late Campanian, being their oldest record of the Weddellian Biogeographic Province.

The skull of a fossil Prion (Aves: Procellariiformes) from the Neogene (Late Miocene) of northern Chile

Se describe un craneo fosil de un procellariido, Pachyptila sp., proveniente de sedimentos marinos del Mioceno Tardio de la Formacion Bahia Inglesa (Mioceno Medio-Plioceno) del norte de Chile. El fosil es comparado con especies actuales de la familia Procellariidae. Este hallazgo representa el primer registro fosil neogeno del genero Pachyptila en America del Sur.

Late Jurassic sauropods in chilean Patagonia

Abstract. A description is provided of the first sauropod remains (i.e., isolated vertebrae and appendicular bones) from the Late Jurassic of Aysén, in Chilean Patagonia (Toqui Formation, late Tithonian). Although the bones found are fragmentary, they still allow the recognition of an unsuspected sauropod diversity for this period in South America. The materials suggest the presence of at least three different sauropod lineages: an indeterminate group of sauropods, possible Titanosauriformes, and Diplodocoidea. A phylogenetic analysis of this last clade supports the placement of the remains within Diplodocinae and also provides the first unequivocal record of this clade in Late Jurassic rocks of South America. These records provide important information about the poorly known evolutionary history of sauropods in South America before the Cretaceous.

Forelimb Posture in Chilesaurus diegosuarezi (Dinosauria, Theropoda) and Its Behavioral and Phylogenetic Implications

Abstract. Many dinosaur skeletons show evidence of behavior, including feeding, predation, nesting, and parental care. The resting posture of the forelimbs has been studied in some theropod species, in relation to the acquisition of flight in advanced maniraptoran theropods. Chilesaurus diegosuarezi is a bizarre tetanuran recently described from the Toqui Formation (latest Tithonian) of southern Chile that is represented by multiple well-preserved and articulated specimens. The aim of the present work is to analyze the forelimb posture of four articulated specimens of Chilesaurus diegosuarezi, focusing on its anatomical description, and phylogenetic and behavioral implications. All the preserved specimens show strongly ventrally flexed arms with the hands oriented backwards, an arrangement that closely resembles those in dinosaur specimens previously described as preserving resting posture, such as Mei long, Sinornithoides youngi and Albinykus baatar. As a result, it seems that individuals of Chilesaurus diegosuarezi have been in passive activity (e.g. feeding, resting) when they were buried quickly, allowing their fossilization in life position and preserving the forelimb resting posture. The arrangement of the forelimb bones in Chilesaurus could show the first evidences of the structures linked to the muscles that flex the forearms, features related with the acquisition of flying control in advanced maniraptorans.