Emmet J. Judziewicz

A worldwide phylogenetic classification of the Poaceae (Gramineae)

Based on recent molecular and morphological studies we present a modern worldwide phylogenetic classification of the ± 12074 grasses and place the 771 grass genera into 12 subfamilies (Anomochlooideae, Aristidoideae, Arundinoideae, Bambusoideae, Chloridoideae, Danthonioideae, Micraioideae, Oryzoideae, Panicoideae, Pharoideae, Puelioideae, and Pooideae), 6 supertribes (Andropogonodae, Arundinarodae, Bambusodae, Panicodae, Poodae, Triticodae), 51 tribes (Ampelodesmeae, Andropogoneae, Anomochloeae, Aristideae, Arundinarieae, Arundineae, Arundinelleae, Atractocarpeae, Bambuseae, Brachyelytreae, Brachypodieae, Bromeae, Brylkinieae, Centotheceae, Centropodieae, Chasmanthieae, Cynodonteae, Cyperochloeae, Danthonieae, Diarrheneae, Ehrharteae, Eragrostideae, Eriachneae, Guaduellieae, Gynerieae, Hubbardieae, Isachneae, Littledaleeae, Lygeeae, Meliceae, Micraireae, Molinieae, Nardeae, Olyreae, Oryzeae, Paniceae, Paspaleae, Phaenospermateae, Phareae, Phyllorachideae, Poeae, Steyermarkochloeae, Stipeae, Streptochaeteae, Streptogyneae, Thysanolaeneae, Triraphideae, Tristachyideae, Triticeae, Zeugiteae, and Zoysieae), and 80 subtribes (Aeluropodinae, Agrostidinae, Airinae, Ammochloinae, Andropogoninae, Anthephorinae, Anthistiriinae, Anthoxanthinae, Arthraxoninae, Arthropogoninae, Arthrostylidiinae, Arundinariinae, Aveninae, Bambusinae, Boivinellinae, Boutelouinae, Brizinae, Buergersiochloinae, Calothecinae, Cenchrinae, Chionachninae, Chusqueinae, Coicinae, Coleanthinae, Cotteinae, Cteniinae, Cynosurinae, Dactylidinae, Dichantheliinae, Dimeriinae, Duthieinae, Eleusininae, Eragrostidinae, Farragininae, Germainiinae, Gouiniinae, Guaduinae, Gymnopogoninae, Hickeliinae, Hilariinae, Holcinae, Hordeinae, Ischaeminae, Loliinae, Melinidinae, Melocanninae, Miliinae, Monanthochloinae, Muhlenbergiinae, Neurachninae, Olyrinae, Orcuttiinae, Oryzinae, Otachyriinae, Panicinae, Pappophorinae, Parapholiinae, Parianinae, Paspalinae, Perotidinae, Phalaridinae, Poinae, Racemobambosinae, Rottboelliinae, Saccharinae, Scleropogoninae, Scolochloinae, Sesleriinae, Sorghinae, Sporobolinae, Torreyochloinae, Traginae, Trichoneurinae, Triodiinae, Tripogoninae, Tripsacinae, Triticinae, Unioliinae, Zizaniinae, and Zoysiinae). In addition, we include a radial tree illustrating the hierarchical relationships among the subtribes, tribes, and subfamilies. We use the subfamilial name, Oryzoideae, over Ehrhartoideae because the latter was initially published as a misplaced rank, and we circumscribe Molinieae to include 13 Arundinoideae genera. The subtribe Calothecinae is newly described and the tribe Littledaleeae is new at that rank.

The South American Species of Arthrostylidium (Poaceae: Bambusoideae: Bambuseae)

A summary of the 11 species of Arthrostylidium occurring in South America is pre- sented, including descriptions and illustrations of four new species. Two montane species from Colombia and Ecuador, A. ecuadorense and A. youngianum, appear to be related to A. venezuelae and A. pubescens respectively, but differ in details of culm leaf, foliage leaf, inflorescence, and spikelet morphology. Two new taxa from lowland Amazonian Brazil are somewhat atypical of the genus but appear to be best referred to it for now: A. fimbrinodum, with an unusual branching pattern, fimbriate culm nodes, short, few-flowered inflorescences, and small spikelets; and A. grandifolium, with stout, nearly solid culms, very large foliage leaves with elongate inner ligules, and numerous, elongate inflorescences.

Endemic herbaceous bamboo genera of Cuba (Poaceae: Bambusoideae: Olyreae)

A morphological, anatomical, and taxonomic study of the endemic herbaceous bamboo (Poaceae: Bambusoideae: Olyreae) genera Ekmanochloa and Mniochloa (including its new segregate Piresiella) is presented Piresiella strephioides is proposed. An unusual feature of all three genera is the presence of inflorescences consisting of paired, conjugate racemes. Illustrations of the leaf blade anatomy show unusual features such as the absence of fusoid cells in Ekmanochloa and Mniochloa and the presence of compound epidermal papillae in Ekmanochloa. three genera are compared with possible close relatives within the Olyreae

Toliara (Poaceae, Chloridoideae, Cynodonteae), a new grass genus endemic to southern Madagascar

Judziewicz E. L. 2009. — Toliara (Poaceae, Chloridoideae, Cynodonteae), a new grass genus endemic to southern Madagascar. Adansonia, sér. 3, 31 (2): 273-277. ABSTRACT Toliara arenacea Judziewicz (Poaceae), a new genus and species endemic to coastal sand dunes in southwestern Madagascar, is described and illustrated. It is related to the widespread paleotropical genus Perotis Aton (Chloridoideae, Cynodonteae), but differs in its spikelets with awnless glumes. Known only from the region of the type locality near the city of Toliara, it is critically endangered.

Olyra davidseana (Poaceae: Bambusoideae: Olyreae), a New Species from Brazil

ABsTRAcr. Olyra davidseana, a new species from eastern Amazonian Brazil in the 0. glaberrima group, is described, illustrated, and compared to related species in a key. The new species is distinctive in its combination of female spikelets with attenuate glumes; smooth, shiny, female lemmas pubescent with flattened macrohairs only along the basal adaxial margins; obtuse to subcordate leaf blades; and small, congested inflorescences. The Olyreae (Poaceae: Bambusoideae) is a group of 19 genera and 80-90 species native to the American tropics. In most genera in the tribe, the species are defined by vegetative characters, the spikelets generally exhibiting little interspecific variation. The exception to this trend is Olyra L., the largest (23 species), most widespread, and most heterogeneous genus in the tribe. Species of Olyra are differentiated mainly on female floret structure, surface texture, and indument, as shown in the scanning electron micrographs presented in a recent revision (Soderstrom and Zuloaga 1989). Soderstrom and Zuloaga did not attempt an infrageneric classification of Olyra, but it is clear that besides about 10 anomalous species, there are several small, phylogenetically natural groups within the genus, such as: 1) the 0. latifolia group (0. latifolia L., 0. caudata Trin., and possibly 0. buchtienii Hackel), with smooth, shiny, presumably bird-dispersed female florets; 2) the monocarpic 0. ecaudata group (0. ecaudata Doell, 0. standleyi A. Hitchc., and 0. taquara Swallen), with inflorescences borne on separate leafless culms; 3) the 0. fasciculata group (0. fasciculata Trin., 0. obliquifolia Steudel, 0. holttumiana Soderstrom & Zuloaga, and 0. tamanquareana Soderstrom & Zuloaga), with umbel

Six new bamboos (Poaceae: Bambusoideae) from the Venezuelan Guayana

The grass flora of the Guayana Highlands is a rich one and includes many poorly known bamboos (Poaceae: Bambusoideae). Twenty-one genera and 75 species of bambusoid grasses are known from the region, including the following six species newly described for the Flora of the Venezuelan Guayana. Judging from the intriguing sterile but as yet unidentifiable bamboo collections that remain in herbaria, more novelties may be expected from this region in the future.

Packera insulae-regalis (Asteraceae, Senecioneae), a new species endemic to Isle Royale, Michigan, U.S.A.

Packera insulae-regalis (Asteraceae, Senecioneae) is a new species that is only known from one population occurring near the summit of Mount Franklin on Isle Royale, Michigan, U.S.A. It resembles both P. paupercula and P. indecora but is hexaploid, whereas the former has diploid and tetraploid populations and the latter is octoploid around the western Great Lakes. Whereas most species of Packera are self-incompatible (as is P. paupercula) and a few are self-compatible (as is P. indecora), P. insulae-regalis is unique in the genus in being partially self-compatible, as far as is known. Its origin post-glacially by hybridization between these two species is hypothesized.

Aulonemia yanachagensis (Poaceae: Bambusoideae: Bambuseae): A new species from central Peru

Aulonemia yanachagensis is described from cloud forests at 2600–3000 m elevation in the Parque Nacional Yanachaga-Chemillén Prov. Oxapampa, Dept. Pasco, Peru. It differs from all other congeners in its combination of prominent sheath auricles, with abundant radiate fimbriae, small foliage leaf blades, and awnless spikelets.ResumenSe describeAulonemia yanachagensis, la cual crece en el bosque de nieblas del Parque Nacional Yanachaga-Chemillén Prov. Oxampampa, Dept. Pasco, Perú, a elevaciones entre 2600–3000 m. Esta especie se difiere de todos sus congéneres por la combinación de caracteres tales como aurículas prominentes, fimbrias radiales abundantes, láminas foliare pequeñas, y espiguillas sin aristas.

Aulonemia cochabambensis (Poaceae: Bambusoideae: Bambuseae: Arthrostylidiinae), an anomalous new species from Bolivia

Aulonemia cochabambensis (Poaceae: Bambusoideae: Bambuseae: Arthrostylidiinae), a new species from the Department of Cochabamba, Bolivia, is described and illustrated. It has foliage leaves with delicate fimbriae, no sheath auricles, narrow blades, an abaxial dark marginal stripe, and intercostal sclerenchyma; few-flowered paniculate synflorescences; and robust, awned spikelets. The new species is compared with its putative relatives Aulonemia laxa and Arthrostylidium schomburgkii. A key to the species of Aulonemia in Bolivia is also included.ResumenSe describe e ilustra Aulonemia cochabambensis (Poaceae: Bambusoideae: Bambuseae: Arthrostylidiinae), una especie nueva del Departamento de Cochabamba, Bolivia. Se distingue la especie nueva por las hojas de follaje sin aurículas y con fimbrias delicadas y láminas angostas con una raya oscura marginal por el envés; sinflorescencias paniculadas de pocas espiguillas; y espiguillas robustas y aristadas. La especie nueva es comparado con Aulonemia laxa y Arthrostylidium schomburgkii, las especies morfologicamente más relacionadas con A. cochabambensis. Se presenta también una clave a las especies de Aulonemia en Bolivia.

Arthrostylidium berryi (Poaceae, Bambusoideae, Bambuseae, Arthrostylidiinae), a New Species from White Sand Shrublands in Venezuela and Colombia

ABSTRACT Arthrostylidium berryi Judziewicz & Davidse (Poaceae, Bambusoideae, Bambuseae, Arthrostylidiinae), a new species from riversides, seasonally flooded white sand shrublands, and granitic lajas in lowland Venezuela and adjacent Colombia, is described and illustrated. It is easily distinguished from its South American congeners in Arthrostylidium Ruprecht by its strongly reflexed leaf blades and long (4–10 cm) spikelets with numerous (nine to 15) florets.