Francisco J. Vega

EARLY CRETACEOUS ARTHROPODS FROM THE TLAYUA FORMATION AT TEPEXI DE RODRIGUEZ, PUEBLA, MEXICO

The arthropod macrofauna from the Middle Member of the lithographic limestones of the Tlayda Formation, in quarries at Tepexi, Mexico, is comprised of marine and nonmarine components. Marine taxa include a new species of flabelliferid isopod, a new genus and species of an anomuran, and a new genus and species of a brachyuran crab. Remains of an arachnid and an odonate nymph represent nonmarine constituents. Previous paleoenvironmental interpretations of a restricted lagoon, with periodic episodes of marine and freshwater influences are consistent with the nature of the arthropod fauna. Isopod remains, represented only by corpses, that resemble moder ectoparasites of fishes suggest that they are directly associated with the abundant fish remains found in the quarries, either as ectoparasites that released their hosts before they died or possibly as scavengers that fed on fish remains. The next most abundant arthropods are the crabs, most of which are corpses, suggesting that this group lived in or very near to the depositional site of the Tlayia Formation. Based upon the new fossil material, the stratigraphic range for the Aeglidae has been extended to span Albian to Holocene time. Extant representatives of this family inhabit fresh water environments of South America.

NEW CRABS FROM THE EOCENE AND OLIGOCENE OF BAJA CALIFORNIA SUR, MEXICO AND AN ASSESSMENT OF THE EVOLUTIONARY AND PALEOBIOGEOGRAPHIC IMPLICATIONS OF MEXICAN FOSSIL DECAPODS

Abstract A new collection of fossil decapod crustaceans from the Cretaceous Rosario Formation, the Eocene Tepetate Formation and the Oligocene El Cien Formation, Baja California Sur, Mexico, has yielded two new genera and several new species, Amydrocarcinus dantei n. gen. and sp., Levicyclus tepetate n. gen. and sp., Eriosachila bajaensis n. sp., Oregonia spinifera n. sp., Archaeopus mexicanus n. sp., and Necronectes nodosa n. sp. Additionally, new occurrences of the previously described Lophoranina bishopi, Xandaros sternbergi, Icriocarcinus xestos, and Lobonotus mexicanus as well as Dardanus cf. D. mexicanus are reported. As part of ongoing work on global evolutionary and paleobiogeographic patterns within the Decapoda, the work has prompted a review and synthesis of decapod occurrences in the tropical and subtropical Americas including the southern United States, the Caribbean, Mexico, Central America, and northern South America. As a result of the systematic review, several new combinations are reported herein which include Eriosachila bartholomaeensis (Rathbun, 1919), Lobonotus sandersi (Blow and Manning, 1996; 1998), and Matutites americanus (Rathbun, 1935). Icriocarcinus is transferred to the Goneplacidae, extending the range of that family into the Cretaceous. Most Cretaceous through Miocene tropical and subtropical American taxa appear to have originated within the area and a large number were endemic. Most of the immigrants to the central Americas appear to have evolved along North Atlantic shelves and subsequently dispersed to the Americas, probably via continental shelf routes. In addition, as demonstrated by several previous studies, decapod crustaceans appear to have evolved in numerous middle- and high- latitude areas with subsequent dispersal to lower latitudes, contrary to the long held notion that the tropics are areas of origin with subsequent dispersal to other regions. Low-latitude decapod taxa tend to remain in low-latitude areas. The Maastrichtian and the Eocene appear to have been times of elevated extinctions within the Decapoda; however, the extinction patterns for those two time intervals are very complex.

Chicxulub impact ejecta from Albion Island, Belize

Impact ejecta from the Albion Formation are exposed in northern Belize. The ejecta come from the outer portion of the continuous ejecta blanket of the Chicxulub crater, which is located 360 km to the northwest. The basal unit of the Albion Formation is a ∼1-m-thick clay and dolomite spheroid bed composed of up to four discrete flows. The clay spheroids are altered impact glass, and the dolomite spheroids are accretionary lapilli. The upper unit is a ∼15-m-thick coarse diamictite bed containing altered glass, large accretionary blocks, striated, polished, and impacted cobbles, and rare shocked quartz. The abundance of accretionary clasts, evidence for atmospheric drag sorting, and the presence of multiple flows in the Albion Formation indicate that atmospheres play an important role in the formation of the outer portions of continuous ejecta blankets of large craters.

Provenance of Upper Cretaceous–Paleogene sandstones in the foreland basin system of the Sierra Madre Oriental, northeastern Mexico, and its bearing on fluvial dispersal systems of the Mexican Laramide Province

Sandstone petrography and detrital zircon U-Pb analysis of Upper Cretaceous–Paleogene sandstones in the foreland basin of the Sierra Madre Oriental in northeastern Mexico indicate long-distance sediment transport from arc, basement, and thrust-belt sources lying to the west, northwest, and south. The basin fill, termed the Difunta Group, consists of sublitharenites, litharenites, feldspathic litharenites, and lithic arkoses derived from mixed sources that included sedimentary rocks, magmatic arc rocks, and subordinate basement rocks. Six age populations comprise the detrital zircon content of the sandstones: Proterozoic (1900–900 Ma), early Paleozoic (500–400 Ma), late Paleozoic–Early Triassic (288–235 Ma), Jurassic (180–151 Ma), Early Cretaceous (150–111 Ma), and Late Cretaceous–Paleogene (110–54 Ma). These grains were derived from several arc terranes, ranging in age from Permian to Paleogene, in western Mexico and the southwestern United States, from sedimentary rocks and possibly interbedded tuffs of the Sierra Madre Oriental orogen and from basement sources or their derivative sandstones of the southwestern United States. The petrographic and geochronologic provenance data corroborate existing models for derivation of much foreland detritus from arc sources to the west, identify the Sierra Madre orogen itself as an important source for sediment, and these data modify the Late Cretaceous–Paleogene paleogeography of Mexico to include a long, orogen-parallel fluvial system with headwaters in the southwestern United States. The difference in average ages of the youngest grains in the sandstones and their inferred depositional ages is 10.5 m.y., indicating that the initial coarse fill of the foreland basin was derived from early Laramide uplift and eastward arc migration in northwestern Mexico and the southwestern United States.

Basinward transport of Chicxulub ejecta by tsunami-induced backflow, La Popa basin, northeastern Mexico, and its implications for distribution of impact-related deposits flanking the Gulf of Mexico

Ejecta-bearing strata are present at the top of Cretaceous foreland-basin deposits throughout the La Popa basin in northeastern Mexico. In the southeast part of the basin, locally thick (as much as 4.6 m) ejecta-rich conglomeratic strata occupy valley-like features at a bathymetric break that separated Maastrichtian upper shoreface from lower shoreface and prodelta depositional settings. Clast-supported textures, normally graded planar conglomerate-sandstone couplets, upcurrent-dipping low-angle cross-laminae, sparse paleocurrent data, and transported fossils indicate deposition by south- to southeast-directed turbulent, supercritical flow. In the northwest part of the basin, ejecta grains are present but less common in correlative deposits. Sediment, ejecta, and organisms were eroded from shoreward environments and transported basinward by backflow of run-up surge(s) emplaced against the continent by one or several tsunami(s). High-discharge, supercritical offshore-directed flow provides a mechanism for transport of voluminous, ejecta-bearing sediment and late Maastrichtian marine organisms into deep-water Gulf of Mexico settings.

Maastrichtian crustacea (Brachyura : Decapoda) from the Ocozocuautla Formation in Chiapas, southeast Mexico

Abstract More than thirty complete specimens of Carcineretes planetarius Vega, Feldmann, Ocampo, and Pope, 1997, a member of the extinct decapod family Carcineretidae, have been collected from the upper part of the Ocozocuautla Formation in Chiapas, southeast Mexico. Stratigraphic occurrences of Carcineretes in the Caribbean Province suggest that this crab should be regarded as an index fossil for the early Maastrichtian. Six samples of this species may represent individuals that died during molting. The sudden disappearance of this family at the end of the Maastrichtian and its restricted paleobiogeographic distribution in the vicinity of the impact site suggest that the Carcineretidae may have been affected by the Chicxulub impact. Other decapod specimens collected from the same localities were assigned to the Xanthidae; Parazanthopsis meyapaquensis new genus and species, and Megaxantho zoque, new genus and species, are described. They constitute the second and third reports of Cretaceous xanthid crabs from Mexico. A lagoonal paleoenvironment is suggested, based on associated fauna and flora. Occurrences of index species of benthic and planktic foraminifera along with that of diagnostic rudist species confirm an early Maastrichtian age.

NEW MIDDLE EOCENE DECAPODS (CRUSTACEA) FROM CHIAPAS, MEXICO

Abstract Decapod crustacean specimens from the middle Eocene San Juan Formation in central Chiapas represent the first record of Eocene decapods in southern México. New taxa include: Dardanus mexicanus new species (Diogenidae), Lophoranina cristaspina new species, Notopus minutus new species (Raninidae); Verrucoides stenohedra new genus and new species (Xanthidae); Stoaplax nandachare new genus and new species (Goneplacidae); and Viapinnixa alvarezi new species (Pinnotheridae). Verrucoides verrucoides new genus and new combination from the Paleocene of Greenland represents a new combination. In addition, the fauna includes Callianassa sensu lato sp., Laeviranina sp., Calappilia cf. C. hondoensis Rathbun, 1930, Eriosachila sp., and indeterminate calappid and xanthoid taxa. This assemblage bears close relationship with coeval faunas in the Tethyan region of southern Europe and southern North America and with Paleocene faunas of Greenland, strengthening the evidence for previously described patterns of dispersal within the Decapoda.

A new species of Late Cretaceous crab (Brachyura: Carcineretidae) from Albion Island, Belize).

A new species of carcineretid crab, Carcineretes planetarius, is described from the Upper Cretaceous (lower Maastrichtian) Barton Creek Dolomite at Albion Island, Belize. The age is based on the stratigraphic range of associated nerineid gastropods and correlation with nannoplankton, benthic foraminifera, and the other known congeneric species of crab found in Jamaica. Confirmation of this age aids in constraining the timing of ejecta deposits of the Chicxulub impact found at the top of Barton Creek Dolomite exposed on Albion Island. Paleoenvironmental and paleoecological analyses suggest that these crabs were swimmers in lagoonal settings, capable of burrowing a few centimeters into the mud for protection.

Isotopic composition of low-latitude paleoprecipitation during the Early Cretaceous

The response of the hydrologic cycle in global greenhouse conditions is important to our understanding of future climate change and to the calibration of global climate models. Past greenhouse conditions, such as those of the Cretaceous, can be used to provide empirical data with which to evaluate climate models. Recent empirical studies have utilized pedogenic carbonates to estimate the isotopic composition of meteoric waters and calculate precipitation rates for the AptianAlbian. These studies were limited to data from mid- (35°N) to high (75°N) paleolatitudes, and thus future improvements in accuracy will require more estimates of meteoric water compositions from numerous localities around the globe. This study provides data for tropical latitudes (18.5°N paleolatitude) from the Tlayua Formation, Puebla, Mexico. In addition, the study confi rms a shallow nearshore depositional environment for the Tlayua Formation. Petrographic observations of fenestral fabrics, gypsum crystal molds, stromatolitic structures, and pedogenic matrix birefringence fabric support the interpretation that the strata represent deposition in a tidal fl at environment. Carbonate isotopic data from limestones of the Tlayua Formation provide evidence of early meteoric diagenesis in the form of meteoric calcite lines. These trends in δ 18 O versus δ 13 C

Chicxulub impact ejecta deposits in southern Quintana Roo, México, and central Belize

Discoveries of Chicxulub impact ejecta of the Albion Formation in road cuts and quarries in southern Quintana Roo, Mexico and Belize, broaden our understanding of ejecta depositional processes in large impacts. There are numerous new exposures of ejecta near the Rio Hondo in Quintana Roo Mexico, located at distances of 330– 350 km from the center of the Chicxulub crater. A single ejecta exposure was discovered near Armenia in central Belize, 470 km from Chicxulub. The Albion Formation is composed of two lithostratigraphic units: the spheroid bed and diamictite bed, originally identifi ed at Albion Island, Belize. The new spheroid bed exposures range from