Jorge O. Ares

Above-ground phytomass dynamics in a grassland steppe of Patagonia, Argentina.

Aerial phytomass and litter dynamics of a grassland steppe in Patagonia, Argentina, were studied at either monthly or bimonthly intervals for 2 years. This area is characterized by a cold and wet winter (June to September) and a warm and dry summer (December to March). The growing season extends from September to April, with moist spring and fall periods interrupted by a midsummer drought. Festuca pallescens (St. Yves) Parodi produced about 95% of the total annual phytomass. Peaks of green phytomass were recorded in spring-summer of the first growing season (33.6 +/- 2.9 g . m-2) and in early spring (35.0 +/- 2.4 g . m-2) and fall (32.7 +/- 2.7 g . m-2) of the second growing season. Less abundant forage grasses were Poa ligularis Nees ap Steudel, Bromus setifolius Presl., Hordeum comosum Presl., and Rytidosperma virescens (Desvaux) Nicors. Shrubs and forbs represented less than 2% of the total annual phytomass of this grassland. The relationships between phytomass production of the main species and some environmental variables are discussed. These results contribute to the knowledge of above-ground phytomass dynamics and forage availability throughout seasons of this grassland in Patagonia, and are part of the data necessary for proper range management planning. This information will permit the designing of appropriate grazing schedules by balancing the number of grazing animals to the forage resources available.

Sex-related spatial patterns of Poa ligularis in relation to shrub patch occurrence in northern Patagonia

Poa ligularis is a dioecious species and a valuable forage plant which is widespread in the arid steppe of northern Patagonia (Argentina). The vegetation in these areas consists of a system of perennial plant patches alternating with bare soil areas defining contrasting micro-environments. We hy- pothesized that (1) male and female individuals of P. ligularis are spatially segregated in different micro-environments, (2) the intensity of spatial segregation of sexes depends on plant structure and (3) spatial segregation of sexes is enhanced by competitive interactions between the sexes within the veg- etation patches. We analysed the spatial distribution of fe- male and male individuals in relation to the spatial pattern of vegetation in two areas differing in their vegetation struc- ture. The location of P. ligularis within patches where either male, female or both sexes occurred was also ana- lysed. The results indicate that different patterns of spatial distribution of sexes of P. ligularis may be found at the community level depending on the dominant life forms and geometric structure of plant patches. Where patches are of a lower height, with a high internal patch cover, individuals of both sexes are concentrated within patch canopies. In sites characterized by large, tall patches and less internal patch cover suitable microsites for female and male P. ligularis occur both within and outside the patch with males located at further distances from the patch edge. Where the patch is large and tall enough to allow the estab- lishment of males and females at relatively high numbers, males occupy the patch periphery or even colonize the inter- patch bare soil. These spatial patterns are consistent with selective traits in which females better tolerate intraspecific competition than males, while males tolerate wider fluctua- tions in the physical environment (soil moisture, nitrogen availability, wind intensity, etc.).

Phenology of Festuca pallescens in relation to topography in north-western Patagonia

. The phenological changes in populations of Festuca pallescens (St. Yves) Parodi at different topographic positions and exposure along an altitudinal gradient (600 - 1100 m) were investigated during two growing seasons in northwestern Patagonia. Stepwise multiple regression analysis was used to describe the relationship between phenology and environment during the entire growing season. Analysis of variance was also performed at each sample date to detect significant environmental factors influencing phenology at different sites. The sum of maximum air temperatures was identified as the environmental variable best correlated with the seasonal variation of phenological events of Festuca pallescens over the period of two growing seasons, explaining 93.2 % of the total variance. Significant differences between sites were observed at each sample date. Main effects of altitude and topographic position and two-way interactions between altitude and topographic position, and topographic position and exposure were also detected as significant. Phenology was delayed at increased altitude. Differences in phenology between topographic sites at the same altitude were not detected during the entire growing season and were only observed in the reproductive phase. At this time, the phenology was significantly delayed at high topographic positions on the slopes as compared with low and mid positions. At high altitudes in the valley (950 m a. s. 1.), where steep slopes and humid conditions prevail, phenology was delayed on western exposures and low positions. The results adequately summarize and quantify the effect of spatial and temporal environmental variation on the phenological development of Festuca pallescens in northwestern Patagonia.