Jose Miguel Rodriguez Espinosa

TORUS AND ACTIVE GALACTIC NUCLEUS PROPERTIES OF NEARBY SEYFERT GALAXIES: RESULTS FROM FITTING INFRARED SPECTRAL ENERGY DISTRIBUTIONS AND SPECTROSCOPY

We used the CLUMPY torus models and a Bayesian approach to fit the infrared spectral energy distributions and ground-based high angular resolution mid-infrared spectroscopy of 13 nearby Seyfert galaxies. This allowed us to put tight constraints on torus model parameters such as the viewing angle i, the radial thickness of the torus Y, the angular size of the cloud distribution σtorus, and the average number of clouds along radial equatorial rays N 0. We found that the viewing angle i is not the only parameter controlling the classification of a galaxy into type 1 or type 2. In principle, type 2s could be viewed at any viewing angle i as long as there is one cloud along the line of sight. A more relevant quantity for clumpy media is the probability for an active galactic nucleus (AGN) photon to escape unabsorbed. In our sample, type 1s have relatively high escape probabilities, P esc ~ 12%-44%, while type 2s, as expected, tend to have very low escape probabilities. Our fits also confirmed that the tori of Seyfert galaxies are compact with torus model radii in the range 1-6 pc. The scaling of the models to the data also provided the AGN bolometric luminosities L bol(AGN), which were found to be in good agreement with estimates from the literature. When we combined our sample of Seyfert galaxies with a sample of PG quasars from the literature to span a range of L bol(AGN) ~ 1043-1047 erg s–1, we found plausible evidence of the receding torus. That is, there is a tendency for the torus geometrical covering factor to be lower (f 2 ~ 0.1-0.3) at high AGN luminosities than at low AGN luminosities (f 2 ~ 0.9-1 at ~1043-1044 erg s–1). This is because at low AGN luminosities the tori appear to have wider angular sizes (larger σtorus) and more clouds along radial equatorial rays. We cannot, however, rule out the possibility that this is due to contamination by extended dust structures not associated with the dusty torus at low AGN luminosities, since most of these in our sample are hosted in highly inclined galaxies.

Functional Interaction between Two Transcription Factors Involved in the Developmental Regulation of a Small Heat Stress Protein Gene Promoter

Hahsp17.6G1 is the promoter of a small heat stress protein (sHSP) from sunflower (Helianthus annuus) that is activated during zygotic embryogenesis, but which does not respond to heat stress. We report here the cloning of a transcription factor (TF), sunflower drought-responsive element binding factor 2 (HaDREB2), by one-hybrid interaction with functional cis-elements in Hahsp17.6G1. We have analyzed the functional interaction between HaDREB2 and a second transcription factor, sunflower heat stress factor A9 (HaHSFA9), which was previously assigned to the regulation of Hahsp17.6G1. HaDREB2 and HaHSFA9 synergistically trans-activate the Hahsp17.6G1 promoter in bombarded sunflower embryos. This synergistic interaction is heat stress factor (HSF) specific and requires the binding of both factors to the promoter. The C-terminal region of HaHSFA9 is sufficient for the HSF specificity. Our results represent an example of a functional interaction between members of the Apetala 2 (HaDREB2) and HSF (HaHSFA9) families of transcription factors. We suggest new roles in zygotic embryogenesis for specific members of the AP2 transcription factor family.

The HaDREB2 transcription factor enhances basal thermotolerance and longevity of seeds through functional interaction with HaHSFA9

BackgroundTranscription factor HaDREB2 was identified in sunflower (Helianthus annuus L.) as a drought-responsive element-binding factor 2 (DREB2) with unique properties. HaDREB2 and the sunflower Heat Shock Factor A9 (HaHSFA9) co-activated the Hahsp17.6G1 promoter in sunflower embryos. Both factors could be involved in transcriptional co-activation of additional small heat stress protein (sHSP) promoters, and thus contribute to the HaHSFA9-mediated enhancement of longevity and basal thermotolerance of seeds.ResultsWe found that overexpression of HaDREB2 in seeds did not enhance longevity. This was deduced from assays of basal thermotolerance and controlled seed-deterioration, which were performed with transgenic tobacco. Furthermore, the constitutive overexpression of HaDREB2 did not increase thermotolerance in seedlings or result in the accumulation of HSPs at normal growth temperatures. In contrast, when HaDREB2 and HaHSFA9 were conjointly overexpressed in seeds, we observed positive effects on seed longevity, beyond those observed with overexpression of HaHSFA9 alone. Such additional effects are accompanied by a subtle enhancement of the accumulation of subsets of sHSPs belonging to the CI and CII cytosolic classes.ConclusionOur results reveal the functional interdependency of HaDREB2 and HaHSFA9 in seeds. HaDREB2 differs from other previously characterized DREB2 factors in plants in terms of its unique functional interaction with the seed-specific HaHSFA9 factor. No functional interaction between HaDREB2 and HaHSFA9 was observed when both factors were conjointly overexpressed in vegetative tissues. We therefore suggest that additional, seed-specific factors, or protein modifications, could be required for the functional interaction between HaDREB2 and HaHSFA9.

Non-correlation between atmospheric extinction coefficient and TOMS aerosol index at the Canarian Observatories

Satellite data measuring aerosols over the Canary Islands need to be correctly interpreted in accordance with the spatial resolution and spectroscopic channels used. In situ data are still a necessary reference for calibrating and interpreting the aerosol index provided by different spectrometers onboard satellites. A comparison of both techniques (in situ and remote) is discussed in this paper, showing there is no linear correlation between the aerosol index and the extinction coefficient. Here, we present an interpretation of different situations arising and discuss critically the correct interpretation of the aerosol index and images provided by the TOMS in accordance with the dust presence over the Observatories. The aerosol index measured at sectors centered at both Observatories (Observatorio del Roque de los Muchachos -ORM-, on La Palma and Observatorio del Teide -OT-, on Tenerife) is also compared.

Comparison of the aerosol index from satellites and the atmospheric extinction coefficient above the Canarian Observatories

Satellite data measuring aerosols over the Canary Islands need to be correctly interpreted in accordance with the spatial resolution and spectroscopic channels used. In situ data are still a necessary reference for calibrating and interpreting the aerosol index provided by different spectrometers onboard satellites. Most of the airmass flux component arriving at the Canarian Archipelago comes from the North Atlantic Ocean and consists of sea aerosols, i.e. absorbent chloride in the UV which does not affect the extinction in the visible range. African dust intrusions affect the western and eastern Canary Islands differently. Moreover, the presence of a stable inversion layer and the sharp orography of the western islands (La Palma at Tenerife) produce different mass flux patterns in the low (mixing) layers closer to the sea and in the median-upper (or free) troposphere layer. The aerosol index provided by the TOMS (Total Ozone Mapping Spectrometer) is one of the most widely accepted products to detect the daily aerosol content. On the other hand, several techniques have been developed in situ to characterize the presence of dust locally at the Canarian Observatories. In particular, a parameter related to sky transparency, the atmospheric extinction coefficient, has been measured at the Roque de los Muchachos Observatory (ORM) on La Palma since 1984 by the Carslberg Automatic Meridian Circle Telescope (CAMC). In situ aerosols are also available for the Teide Observatory (OT) on Tenerife, from 1986 onwards. In this work we compare aerosol index data of TOMS/Earth Probe observations and atmospheric optical extinction coefficient from CAMC in the period 1996-2004. A preliminary comparison of both techniques (in situ and remote) is discussed in this paper, showing there is no linear correlation between the aerosol index and the extinction coefficient. Here, we present an interpretation of different situations arising and discuss critically the correct interpretation of the aerosol index and images provided by the TOMS in accordance with the dust presence over the Observatories.

Constraining the epoch of reionization from the observed properties of the high-z Universe

We combine observational data on a dozen independent cosmic properties at high-

The GTC: a convenient test bench for ELT demonstrations

z

Facility instruments for the GTC

with the information on reionization drawn from the spectra of distant luminous sources and the cosmic microwave background (CMB) to constrain the interconnected evolution of galaxies and the intergalactic medium since the dark ages. The only acceptable solutions are concentrated in two narrow sets. In one of them reionization proceeds in two phases: a first one driven by Population III stars, completed at

New Ground Based Facilities in QSO Research

z\sim 10

Anisotropy in the Mid-IR Emission of Seyfert Galaxies

, and after a short recombination period a second one driven by normal galaxies, completed at