Joseph A. Frederickson

Craniofacial ontogeny in Centrosaurus apertus

Centrosaurus apertus, a large bodied ceratopsid from the Late Cretaceous of North America, is one of the most common fossils recovered from the Belly River Group. This fossil record shows a wide diversity in morphology and size, with specimens ranging from putative juveniles to fully-grown individuals. The goal of this study was to reconstruct the ontogenetic changes that occur in the craniofacial skeleton of C. apertus through a quantitative cladistic analysis. Forty-seven cranial specimens were independently coded in separate data matrices for 80 hypothetical multistate growth characters and 130 hypothetical binary growth characters. Both analyses yielded the max-limit of 100,000 most parsimonious saved trees and the strict consensus collapsed into large polytomies. In order to reduce conflict resulting from missing data, fragmentary individuals were removed and the analyses were rerun. Among both the complete and the reduced data sets the multistate analyses recovered a shorter tree with a higher consistency index (CI) than the additive binary data sets. The arrangement within the trees shows a progression of specimens with a recurved nasal horn in the least mature individuals, followed by specimens with straight nasal horns in relatively more mature individuals, and finally specimens with procurved nasal horns in the most mature individuals. The most mature individuals are further characterized by the reduction of the cranial horn ornamentations in late growth stages, a trait that similarly occurs in the growth of other dinosaurs. Bone textural changes were found to be sufficient proxies for relative maturity in individuals that have not reached adult size. Additionally, frill length is congruent with relative maturity status and makes an acceptable proxy for ontogenetic status, especially in smaller individuals. In adult-sized individuals, the fusion of the epiparietals and episquamosals and the orientation of the nasal horn are the best indicators of relative maturity. This study recovers no clear evidence for sexually specific display structures or size dimorphism in C. apertus.

New Cretaceous lungfishes (Dipnoi, Ceratodontidae) from western North America

Abstract. Ceratodontid lungfishes are generally rare, poorly represented elements of North Americas Mesozoic ecosystems, with previously known maximum diversity in the Late Jurassic. Herein we describe four new species of the form genus Ceratodus, from the Cretaceous of the Western Interior, considerably expanding fossil representation of post-Triassic dipnoans in North America. To model taxonomic and morphologic diversity, we adopt a four-fold system of phenetically based species groups, named for exemplars from the Morrison Formation. Ceratodus kirklandi n. sp. (Potamoceratodus guentheri group) and C. kempae n. sp. (C. frazieri group) represent a hitherto unsampled time interval, the Valanginian. Ceratodus nirumbee n. sp. and C. molossus n. sp. extend the temporal ranges of the C. fossanovum and C. robustus groups upward to the Albian and Cenomanian, respectively. These new occurrences show that ceratodontids maintained their highest diversity from the Late Jurassic through the mid-Cretaceous (Albian—Cenomanian), an interval of ∼60 Myr. The existing record suggests that some of the later (mid-Cretaceous) ceratodontids may have been tolerant of salt water; to date, there is no evidence that they aestivated. Only a few occurrences are known from horizons younger than Cenomanian. Demise of ceratodontids appears to be part of a broader pattern of turnover that occurred at the Cenomanian-Turonian boundary in North America.

A Gigantic Shark from the Lower Cretaceous Duck Creek Formation of Texas

Three large lamniform shark vertebrae are described from the Lower Cretaceous of Texas. We interpret these fossils as belonging to a single individual with a calculated total body length of 6.3 m. This large individual compares favorably to another shark specimen from the roughly contemporaneous Kiowa Shale of Kansas. Neither specimen was recovered with associated teeth, making confident identification of the species impossible. However, both formations share a similar shark fauna, with Leptostyrax macrorhiza being the largest of the common lamniform sharks. Regardless of its actual identification, this new specimen provides further evidence that large-bodied lamniform sharks had evolved prior to the Late Cretaceous.

A New Species of the Lungfish Ceratodus (Dipnoi) from the Early Cretaceous of the Eastern U.S.A.

http://zoobank.org/urn:lsid:zoobank.org:pub:CFECA48B-7833-472B-8FA8-D746B3A6D9CE SUPPLEMENTAL DATA—Supplemental materials are available for this article for free at www.tandfonline.com/UJVP Citation for this article: Frederickson, J. A., T. R. Lipka, and R. L. Cifelli. 2016. A new species of the lungfish Ceratodus (Dipnoi) from the Early Cretaceous of the eastern U.SA. Journal of Vertebrate Paleontology. DOI: 10.1080/02724634.2016.1136316.

First reported actinopterygian from the Navajo Sandstone (Lower Jurassic, Glen Canyon Group) of southern Utah, USA

Abstract. We report the first occurrence of an actinopterygian fish from the Lower Jurassic Navajo Sandstone, discovered in the Grand Staircase-Escalante National Monument in southern Utah, U.S.A. The site contains multiple individuals, preserved within an interdune deposit, possessing the elongate modified dorsal scales usually characterizing semionotiform fishes. The presence of moderately sized fish provides further evidence that interdune oases were occasionally persistent environmental habitats within the greater Navajo dune system, and that the paleobiota is still woefully undersampled. Additionally, this site could help fill a gap in the actinopterygian fossil record between the patchy Lower Jurassic and better-known Middle Jurassic documentation of western North America.

Faunal composition and paleoenvironment of the Arundel Clay (Potomac Formation; Early Cretaceous), Maryland, USA

The Arundel Clay facies of the Potomac Group represents one of the few Lower Cretaceous vertebrate-bearing deposits in the Atlantic coastal plain. Vertebrate fossils from this unit have been known for more than 150 years, but thus far formal descriptions have mainly concentrated on its dinosaurs and mammals. Herein, we eport on a moderately diverse faunal assemblage (USNM 41614) from Dinosaur Park in Prince Georges County, Maryland. This assemblage is represented by 306 disarticulated macroand microfossils that largely consist of teeth and scales (89%). This vertebrate fauna includes two species of hybodont sharks, multiple semionotid fishes, one species of lungfish, three species of turtle, three families of neosuchian crocodilians, six species of dinosaurs, and two species of mammals. Combined with other historical collections from this unit, these new additions to the fauna show that the Arundel was a far more robust and diverse ecosystem than previously envisaged, broadly similar in composition to contemporaneous units of western North America. The Arundel assemblage differs, however, from those in many other Lower Cretaceous sites in that it is dominated numerically by Hybodus and goniopholidid crocodylomorphs, which together comprise 58% of catalogued specimens. Similarly, this sample entirely lacks lissamphibians and lepidosaurs. Traditionally, the Arundel has been interpreted as being of fluvial origin, deposited in a freshwater system of stranded channels or oxbows. Based on faunal composition, together with published geological and sedimentological evidence, we propose that at least some of the Arundel facies was deposited in close proximity to the Atlantic Ocean. Joseph A. Frederickson. Sam Noble Museum, 2401 Chautauqua Avenue, Norman, Oklahoma, 73072, USA and Department of Biology, University of Oklahoma, Norman, Oklahoma, 73019, USA. joe.frederickson@swosu.edu Thomas R. Lipka. Sam Noble Museum, 2401 Chautauqua Avenue, Norman, Oklahoma, 73072, USA thomas.r.lipka-1@ou.edu Richard L. Cifelli. Sam Noble Museum, 2401 Chautauqua Avenue, Norman, Oklahoma, 73072, USA and Department of Biology, University of Oklahoma, Norman, Oklahoma, 73019, USA. rlc@ou.edu