Juan José Ponce

TAPHONOMY AND PALEOECOLOGY OF THE BIVALVE TRACE FOSSIL PROTOVIRGULARIA IN DELTAIC HETEROLITHIC FACIES OF THE MIOCENE CHENQUE FORMATION, PATAGONIA, ARGENTINA

Abstract Lower Miocene tide-influenced deltaic deposits from the Chenque Formation, Patagonia, Argentina, contain abundant and well-preserved biogenic structures attributed to locomotion of deposit-feeder protobranch bivalves. These trace fossils, assigned to the ichnogenus Protovirgularia, consist of delicate, inclined-to-horizontal, chevronate structures, mostly symmetrical with respect to a median axis. Identification of Protovirgularia at sandstone sole beds (hypichnion) is quite straightforward. Endichnial, exichnial and epichnial preservation in heterolithic facies, however, provides a wide variety of forms that depart from the archetypal Protovirgularia and challenges ichnotaxonomic classification. Specimens in prodelta and delta-front facies display morphologic features controlled by substrate fluidity, toponomy, and sedimentation rate. Most specimens show sharp, closely spaced chevrons and occur along sandstone/mudstone interfaces of the proximal prodelta and distal delta-front deposits. These forms reflect how tracemakers experienced significant friction while advancing through the sediment, which resulted in relatively smaller increments of movements. In contrast, variants of Protovirgularia formed in muddier beds, such as in prodeltaic facies, show irregular, poorly defined and unevenly spaced chevrons, and are locally asymmetric in relation to the axis, reflecting softer, water-rich, and plastic substrates. This sediment offered relatively low friction but poor anchorage for the foot. These occurrences of Protovirgularia in tide-influenced, marginal-marine deposits suggests that protobranchs were tolerant of fluctuations in salinity, sedimentation rates, turbidity, and oxygen depletion, displaying opportunistic strategies in stressed nearshore environments. Our evaluation of taphonomic controls and appropriate identification of Protovirgularia can provide valuable information for expanding our knowledge of the ethology and paleoecology of protobranch bivalves.

Ichnology of Deltaic Mouth-Bar Systems of the Lajas Formation (Middle Jurassic) in the Sierra de la Vaca Muerta, Neuquén Basin, Argentina

Abstract. Ichnologic and sedimentologic studies of the Lajas Formation (Middle Jurassic) in Sierra de la Vaca Muerta allowed the recognition of two different types of deltaic mouth bars, each of them showing trace fossil suites with different characteristics. Type I deltaic mouth bars consist of fine to coarse sandstones and fine conglomerates completely reworked by fair-weather and storm wave action, revealing a predominance of basinal hydraulic processes (e.g., waves) during bar construction and progradation. Trace fossil assemblages are composed of Ophiomorpha and Haentzschelinia in the foreset beds, and Polykladichnus, Skolithos, and Arenicolites in the topset beds. Type II deltaic mouth bars comprise sandstones that are fine to coarse and massive or present high angle cross-stratification and current ripples migrating in the opposite direction to the inclination of the foresets. These bars are interpreted to have been deposited during intervals of extraordinary fluvial discharge when wave action was restricted to the topset part of the bars. Whereas equilibrium trace fossils occur in the bottomset beds, escape trace fossils and Ophiomorpha are recorded in the distal foreset beds. In the topset beds, Skolithos and Polykladichnus specimens are very abundant. In general, the two types of mouth bars show low diversity, low abundance of trace fossils and a simple tiering structure. Such traits reflect environmental stresses mainly produced by fluctuating hydraulic energy, salinity, sediment input and high mobility of the substrate.

BIOGENIC STRUCTURES OF UNIONIFORM BIVALVES IN WET-INTERDUNE DEPOSITS (LATE MIOCENE–EARLY PLIOCENE, ARGENTINA)

Abstract Environmental changes within a Neogene coastal dune system are recorded by endobenthic unioniform bivalves that lived in muddy or sandy interdune pond sediments. These bivalves were suspension-filter feeders that formed dense, almost monospecific communities in the wet-interdune deposits of the continental intervals of the Río Negro Formation (late Miocene–early Pliocene). Activity of unioniform bivalves appears to be related to sediment type; resting and locomotion traces dominate in the muddy heterolithic facies, whereas equilibrium/escape structures prevail in the sand-dominated heterolithic facies. These traces characterize two scenarios of the wet-interdune development. First, during high and/or relatively stable water levels, bivalves colonized the muddy bottom and produced resting and locomotion traces. When water level dropped due to desiccation, biogenic structures were impacted by the formation of mud cracks and subsequently covered by sand delivered by migrating dunes. Second, in spite of dune migration, some interdune areas remained wet or flooded and, in response to sediment aggradation, the bivalves produced equilibrium or escape structures, depending on the thickness of eolian sand cover. Only the integration of ichnologic and sedimentologic observations allows deciphering the evolution of the Neogene wet-interdune system in such a detail.

Dinoflagellate cyst zonation for the middle to upper Eocene in the Austral Basin, southwestern Atlantic Ocean: implications for regional and global correlation

The well-exposed marine Eocene units from southwestern Patagonia, Argentina, contain useful information for reconstructing regional climate and oceanographic patterns in an area adjacent to the Drake Passage. The aim of this paper is to integrate dinoflagellate cyst data from three sections of the southwestern Austral Basin (Rio Turbio Formation) to propose a zonation scheme, which can be applied to other southwestern Atlantic Ocean sites. Assemblages of organic walled dinoflagellate cysts have been analysed in different cropping-out sections and cores, showing the high potential of this fossil group as biostratigraphic markers. Comparison of dinoflagellate cyst events of the upper member of the Rio Turbio Formation with calibrated biostratigraphic ranges in the Palaeogene South Pacific Ocean allowed us to date and correlate these sedimentary sections. The resulting zonation consists of four dinoflagellate cyst zones labelled RTF 1 to RTF 4, between the middle Lutetian and late Priabonian. As a final point, we applied dinoflagellate cyst species with importance as palaeoenvironmental markers to assess long-term climatic and oceanographic evolution for the area. This study shows that the endemic–Antarctic dinoflagellate cyst assemblage is dominant during the middle to late Eocene (RTF 1 to RTF 3), while a significant replacement of these taxa by cosmopolitan species characterizes the upper part of the upper member of the Rio Turbio Formation (RTF 4). This turnover seems to be a consequence of changes in the ocean circulation patterns forced by deepening of the southern Atlantic gateways (the Drake Passage and the Tasman Gateway).