M. Franco Tortello

Global Cambrian trilobite palaeobiogeography assessed using parsimony analysis of endemicity

Abstract Palaeobiogeographical data on Cambrian trilobites obtained during the twentieth century are combined in this paper to evaluate palaeoceanographic links through c. 30 myr, once these arthropods biomineralized. Worldwide major tectonostratigraphic units are characterized at series intervals of Cambrian time and datasets of trilobite genera (629 for Cambrian Series 2, 965 for Cambrian Series 3, and 866 for the Furongian Series) are analysed using parsimony analysis of endemicity. Special attention is given to the biogeographical observations made in microcontinents and exotic terranes. The same is done for platform-basinal transects of well-known continental margins. The parsimony analysis of endemicity analysis resulted in distinct palaeogeographical area groupings among the tectonostratigraphic units. With these groupings, several palaeobiogeographical units are distinguished, which do not necessarily fit the previously proposed biogeographical realms and provinces. Their development and spatial distributions are broadly controlled by Cambrian palaeoclimates, palaeogeographical conditions (e.g. carbonate productivity and anoxic conditions) and ocean current circulation. Supplementary material: Global dataset of Cambrian Epoch 2 (A), Cambrian Epoch 3 (B) and the Furongian Epoch (C) trilobite genera are provided at: http://www.geolsoc.org.uk/SUP18669

Cambrian agnostoid trilobites from Mendoza, Argentina: A systematic revision and biostratigraphic implications

Agnostoid trilobites from the Carlos Rusconi collections in the Museum of Natural Sciences of Mendoza, Argentina, are revised. The agnostoids were collected in the southern Precordillera of Mendoza Province, western Argentina. Revised taxa include Agnostus microcephalus (Rusconi), Lotagnostus ( Lotagnostus ) peladensis ? (Rusconi), Lotagnostus ( Lotagnostus ) lasherensis (Rusconi), Homagnostus pehuenchensis Rusconi, Kormagnostus seclusus (Walcott), Kormagnostus ? beltensis (Lochman in Lochman and Duncan), Hypagnostus parvifrons (Linnarsson), Ptychagnostus cuyanus (Rusconi), Ptychagnostus aculeatus (Angelin), Lejopyge laevigata (Dalman), Tomagnostella exsculpta (Angelin), Diplagnostus planicauda (Angelin), Oedorhachis typicalis Resser, Clavagnostus calensis Rusconi and Clavagnostus repandus (Westergard in Holm and Westergard). Some taxa of uncertain affinities are recognized and new specimens assignated to Lotagnostus and Pseudagnostinae are illustrated. Hypagnostus parvifrons, L. laevigata and C. repandus are reported for the first time in South America. Global correlation with previously described agnostoid faunas indicates that the outcrops range from middle Middle Cambrian to latest Late Cambrian in age.

EARLY ORDOVICIAN TRILOBITES FROM THE NAZARENO AREA, NORTHWESTERN ARGENTINA

Abstract The upper part of the Santa Rosita Formation (Ordovician, Tremadocian) in the Nazareno area, Cordillera Oriental, northwestern Argentina, records the vertical passage of high-energy, shallow water platform environments to offshore settings. Eighteen trilobite species are described from this locality for the first time. Although the taxa from the lower part of the succession (Leptoplastides sp., Asaphellus sp.) are scarce and non-age diagnostic, those from the upper part include diverse assemblages partially assigned to the late Tremadocian Notopeltis orthometopa Zone. Systematic descriptions of several species (Geragnostus nesossii Harrington and Leanza, G. callaveiformis Harrington and Leanza, Asaphellus jujuanus Harrington, Notopeltis orthometopa [Harrington], Mekynophrys nanna Harrington, Ceratopyge forficuloides Harrington and Leanza, Apatokephalus tibicen Přibyl and Vanĕk) are improved, the genus Nileus Dalman (including N. australis n. sp.) is first reported from the Tremadocian of western Gondwana, and new species of Asaphellus Callaway (A. nazarenensis n. sp.), Conophrys Callaway, and Apatokephalus Brøgger are described. The trilobites have their closest affinities with faunas from Norway and Sweden. Notopeltis orthometopa and Mekynophrys nanna are restricted to the uppermost part of the succession, well above the first records of most other trilobites recognized.

THE EYES OF THE EARLY TREMADOC OLENID TRILOBITE JUJUYASPIS KEIDELI KOBAYASHI, 1936

Abstract The morphology of the eyes of the olenid trilobite Jujuyaspis keideli Kobayashi, 1936, is described and illustrated. Studied specimens come from Early Tremadocian pyritiferous black shales of the Casa Colorada Formation (=Purmamarca shales) at Purmamarca, Jujuy Province, northwestern Argentina. The eyes are holochroal and proportionately large relative to the overall size of cephalon. They are always found attached to the librigena, showing no preserved lenses, only molds of their surfaces. Their molds demonstrate that lenses were numerous, biconvex, hexagonal in outline and arranged in an hexagonal close-packing system. The eye curvature and the disposition of the facets covering all the visual surface indicate that Jujuyaspis keideli Kobayashi had a visual field wider than that of most benthic olenids. The pattern of lens arrangement and the poorly developed peripheral zone support pelturine affinities for the species.

Early Ordovician trilobites from the Iruya area (Cordillera Oriental, northwestern Argentina) and their stratigraphic significance

Abstract. The middle part of the Santa Rosita Formation (Tremadocian) is well exposed in the Iruya area, northwestern Argentina. At the Pantipampa and Rodeo Colorado localities, it is composed of shale and sandstone representing a wave-dominated shelf with influence of storm activity. Twenty-two trilobite species are described from these localities. Low-diversity assemblages from the lower part of the succession (Kainella meridionalis Kobayashi, Asaphellus catamarcensis Kobayashi, Leptoplastides marianus [Hoek]) are diagnostic of the early Tremadocian Kainella meridionalis Zone. Trilobites from the middle part of the sequence are much more diverse. Kainella teiichii Vaccari and Waisfeld, Gymnagnostus kobayashii n. sp., Conophrys sp. indet., Asaphellus clarksoni n. sp., A. stenorhachis (Harrington), A. isabelae Meroi Arcerito, Waisfeld and Balseiro, Ogygiocaris? iruyensis n. sp., Niobe (Niobella) inflecta (Harrington and Leanza) n. comb., Metayuepingia riccardii n. sp., Pseudokainella keideli Harrington, Apatokephalus rugosus n. sp., Onychopyge acenolazai n. sp., O. gonzalezae n. sp., Nileus cingolanii n. sp., N. erici n. sp., Leptoplastides marianus, Parabolinella sp. indet., Hapalopleura sp. indet., and Ceratopygidae gen. et sp. indet., occur at different levels of the Kainella teiichii Zone. This biostratigraphic unit includes the oldest records of Nileus Dalman and Ogygiocaris? Angelin; Metayuepingia Liu, Niobe (Niobella) Reed, and Onychopyge Harrington are here first reported from the Tremadocian of southwest Gondwana. Finally, the uppermost part of the succession is characterized by the absence of the genus Kainella Walcott and the occurrence of Bienvillia tetragonalis (Harrington), Asaphellus stenorhachis, Pseudokainella keideli and Leptoplastides sp. indet., which are indicative of the middle Tremadocian Bienvillia tetragonalis Zone. The trilobites described in this paper provide a basis for the refinement of correlations with other Lower Ordovician sections of the Cordillera Oriental. The genera recognized have their closest affinities with faunas from Scandinavia, Great Britain, and China.