Marco Marzola
Universidade Nova de Lisboa
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Featured researches published by Marco Marzola.
Historical Biology | 2015
Marco Marzola; João Russo; Octávio Mateus
Eggshells from the three extant crocodilian species Crocodylus mindorensis (Philippine Crocodile), Paleosuchus palpebrosus (Cuviers Smooth-fronted Caiman or Musky Caiman) and Alligator mississippiensis (American Alligator or Common Alligator) were prepared for thin section and scanning electron microscope analyses and are described in order to improve the knowledge on crocodilian eggs anatomy and microstructure, and to find new apomorphies that can be used for identification. Both extant and fossil crocodilian eggs present an ornamentation that vary as anastomo-, ramo- or the here newly described rugosocavate type. The angusticaniculate pore system is a shared character for Crocodylomorpha eggshells and some dinosaurian and avian groups. Previously reported signs of incubated crocodilian eggs were found also on our only fertilised and hatched egg. Paleosuchus palpebrosus presents unique organization and morphology of the three eggshell layers, with a relatively thin middle layer characterised by dense and compact tabular microstructure.
Journal of Vertebrate Paleontology | 2018
Federico L. Agnolin; Octávio Mateus; Jesper Milàn; Marco Marzola; Oliver Wings; Jan Schulz Adolfssen; Lars B. Clemmensen
ABSTRACT The fossil record of post-Paleozoic lungfishes in Greenland is currently restricted to a few brief reports of isolated and undetermined tooth plates coming from the uppermost Fleming Fjord Formation (late Norian) in Jameson Land, central East Greenland. Here, we describe Ceratodus tunuensis, sp. nov., a new dipnoan from a thin bed of calcareous lake mudstone from the Ørsted Dal Member of the Fleming Fjord Formation. The Ceratodus fossil record indicates that during the Late Triassic, this genus was restricted to the middle latitudes of the Northern Hemisphere. This record matches previous paleobiogeographical analyses and indicates that terrestrial biota during the Late Triassic was strongly influenced by paleolatitude.
PLOS ONE | 2017
João Russo; Octávio Mateus; Marco Marzola; Ausenda Balbino
The Late Jurassic Lourinhã Formation is known for its abundant remains of dinosaurs, crocodylomorphs and other vertebrates. Among this record are nine localities that have produced either dinosaur embryos, eggs or eggshell fragments. Herein, we describe and identify the first crocodiloid morphotype eggs and eggshells from the Lourinhã Formation, from five occurrences. One clutch from Cambelas, composed of 13 eggs, eggshell fragments from Casal da Rola and Peralta, one crushed egg and eggshells from Paimogo North, and four crushed eggs as well as eggshell fragments from Paimogo South. We observed and confirmed diagnostic morphological characters for crocodiloid eggshells and which are consistent with a crocodylomorph affinity, such as the ellipsoidal shape, wedge-shaped shell units, triangular extinction under cross-polarized light, and tabular ultrastructure. This material is distinctive enough to propose two new ootaxa within the oofamily Krokolithidae, Suchoolithus portucalensis, oogen. and oosp. nov., for the material from Cambelas, the most complete clutch known for crocodiloid eggs, and Krokolithes dinophilus, oosp. nov., for the remaining material. These are the oldest crocodylomorph eggs known, extending the fossil record for this group to the Late Jurassic. Furthermore, except for the clutch from Cambelas, the material was found with theropod eggs and nests, in the other four occurrences, which seem to suggest some form of biological relationship, still unclear at this point.
Journal of Vertebrate Paleontology | 2017
Marco Marzola; Octávio Mateus; Neil H. Shubin; Lars B. Clemmensen
ABSTRACT Cyclotosaurus naraserluki, sp. nov., is a new Late Triassic capitosaurid amphibian from lacustrine deposits in the Fleming Fjord Formation of the Jameson Land Basin in Greenland. It is based on a fairly complete and well-preserved skull associated with two vertebral intercentra. Previously reported as Cyclotosaurus cf. posthumus, C. naraserluki is unique among cyclotosaurs for having the postorbitals embaying the supratemporals posteromedially. The anterior palatal vacuity presents an autapomorphic complete subdivision by a wide medial premaxillary-vomerine bony connection. The parasphenoid projects between the pterygoids and the exoccipitals, preventing a suture between the two, a primitive condition shared with Rhinesuchidae, Eryosuchus, and Kupferzellia. Within Cyclotosaurus, the Greenlandic skull has a distinctive combination of circular choanae (shared with C. ebrachensis, C. posthumus, and C. robustus) and a convex posteromedial margin of the tabulars (also present in C. ebrachensis and C. intermedius). A phylogenetic analysis indicates that C. naraserluki is the sister taxon of the middle Norian C. mordax from southern Germany, with which it shares a pair of premaxillary foramina. Cyclotosaurus is one of the most successful and diverse genera of Late Triassic temnospondyls, with at least eight species reported from middle Carnian to late Norian. Cyclotosaurus naraserluki is the largest amphibian ever reported from Greenland and one of the Late Triassic vertebrates with the highest northern paleolatitude currently known.
Journal of Vertebrate Paleontology, Program and Abstracts, 2014 | 2014
Marco Marzola; Octávio Mateus; Anne S. Schulp; Louis L. Jacobs; Michael J. Polcyn; Vladimir Pervov
Palaeogeography, Palaeoclimatology, Palaeoecology | 2017
Octávio Mateus; Marco Marzola; Anne S. Schulp; Louis L. Jacobs; Michael J. Polcyn; Vladimir Pervov; António Olímpio Gonçalves; Maria Luísa Morais
Comunicações Geológicas | 2014
João Russo; Octávio Mateus; Ausenda Balbino; Marco Marzola
Bulletin of The Geological Society of Denmark | 2018
Marco Marzola; Octávio Mateus; Jesper Milàn; Lars B. Clemmensen
Journal of Vertebrate Paleontology, Program and Abstracts | 2017
Marco Marzola; Octávio Mateus; Jesper Milàn; Lars B. Clemmensen
Abstract book of the XV Encuentro de Jóvenes Investigadores en Paleontología/XV Encontro de Jovenes Investigadores em Paleontologia, Pombal, 428 pp. | 2017
Marco Marzola; Octávio Mateus; Jesper Milàn; Lars B. Clemmensen