Raquel Rabal-Garcés

Bone Accumulation by Leopards in the Late Pleistocene in the Moncayo Massif (Zaragoza, NE Spain)

Eating habits of Panthera pardus are well known. When there are caves in its territory, prey accumulates inside them. This helps to prevent its kill from being stolen by other predators like hyenas. Although the leopard is an accumulator of bones in caves, few studies have been conducted on existing lairs. There are, however, examples of fossil vertebrate sites whose main collecting agent is the leopard. During the Late Pleistocene, the leopard was a common carnivore in European faunal associations. Here we present a new locality of Quaternary mammals with a scarce human presence, the cave of Los Rincones (province of Zaragoza, Spain); we show the leopard to be the main accumulator of the bones in the cave, while there are no interactions between humans and leopards. For this purpose, a taphonomic analysis is performed on different bone-layers of the cave.

Carnivores from Los Rincones, a leopard den in the highest mountain of the Iberian range (Moncayo, Zaragoza, Spain)

Carnivores are a common element in Pleistocene fossil assemblages. However, they are not so abundant in terms of the numbers of remains. Here, we present a palaeontological study of the carnivores from the Late Pleistocene (MIS 3) of Los Rincones (Spain), one of the few deposits accumulated by leopards. One hundred and ten leopard remains have been recovered. This carnivore is not the only inhabitant of the cave, and 175 remains belonging to Ursus arctos have also been recovered, making it one of the sites with the greatest number of brown bear remains in the Iberian Peninsula. The large number of leopard remains has allowed us to make a detailed morphological and biometrical study that has enabled us to classify the remains within the subspecies Panthera pardus spelaea. The European Ice Age leopard inhabited Europe during Upper Pleistocene and it presents some similarities with Panthera uncia. A study of the scarce remains of Canis lupus indicates that this was similar in size to Canis lupus maximus; the scarcity of the remains prevents us from assigning our remains to this subspecies. A study of the brown bear remains indicates that it is similar to other populations in the north of the Iberian Peninsula with this chronology.

Exceptional crocodylomorph biodiversity of "La Cantalera" site (lower Barremian; Lower Cretaceous) in Teruel, Spain

The palaeontological site of La Cantalera in Teruel, Spain (Blesa Formation, Lower Cretaceous) can be characterized as the site with the greatest biodiversity of vertebrates (32 taxa) from the lower Barremian of the Iberian Peninsula. Remains of amphibians, lizards, turtles, crocodiles, pterosaurs, dinosaurs and mammals have been recovered. The most common taxa are crocodylomorphs, and their most abundant remains are small isolated teeth. Six morphotypes have been identified, which may correspond to at least four different sympatric crocodylomorph taxa. This site presents a similar assemblage to other European Lower Cretaceous sites, with morphotypes that may correspond to the families Goniopholididae, Bernissartiidae, Atoposauridae and multiple ziphodont crocodylomorphs. The crocodylomorph assemblage of La Cantalera is characterized by the absence of large individuals and presents a wide variety of dental morphologies adapted to diets ranging from generalist to highly specialized. This rich biodiversity could be due to concentration of vertebrates into restricted flooded areas during dry seasons within a marsh ecosystem. Furthermore, La Cantalera could also be close to a nesting area, which would explain the abundance of small-sized individuals and the presence of fossil crocodiloid eggshells. Eduardo Puértolas-Pascual. Grupo Aragosaurus-IUCA, Departamento de Ciencias de la Tierra, Facultad de Ciencias, Universidad de Zaragoza, Calle Pedro Cerbuna 12, 50009 Zaragoza (Spain). puertola@unizar.es Raquel Rabal-Garcés. Grupo Aragosaurus-IUCA, Departamento de Ciencias de la Tierra, Facultad de Ciencias, Universidad de Zaragoza, Calle Pedro Cerbuna 12, 50009 Zaragoza (Spain) raquelrabal@hotmail.com José Ignacio Canudo. Grupo Aragosaurus-IUCA, Departamento de Ciencias de la Tierra, Facultad de Ciencias, Universidad de Zaragoza, Calle Pedro Cerbuna 12, 50009 Zaragoza (Spain) jicanudo@unizar.es

Pleistocene cave hyenas in the Iberian Peninsula: new insights from Los Aprendices cave (Moncayo, Zaragoza)

A new Pleistocene paleontological site, Los Aprendices, located in the northwestern part of the Iberian Peninsula in the area of the Moncayo (Zaragoza) is presented. The layer with fossil remains has been dated by amino acid racemization to 143.8 ± 38.9 ka (earliest Late Pleistocene or latest Middle Pleistocene). Five mammal species have been identified in the assemblage: Crocuta spelaea (Goldfuss, 1823) Capra pyrenaica (Schinz, 1838), Lagomorpha indet, Arvicolidae indet and Galemys pyrenaicus (Geoffroy, 1811). The remains of C. spelaea represent a mostly complete skeleton in anatomical semi-connection. The hyena specimen represents the most complete skeleton ever recovered in Iberia and one of the most complete remains in Europe. It has been compared anatomically and biometrically with both European cave hyenas and extant spotted hyenas. In addition, a taphonomic study has been carried out in order to understand the origin and preservation of these exceptional remains. The results suggest rapid burial with few scavenging modifications putatively produced by a medium sized carnivore. A review of the Pleistocene Iberian record of Crocuta spp. has been carried out, enabling us to establish one of the earliest records of C. spelaea in the recently discovered Los Aprendices cave, and also showing that the most extensive geographical distribution of this species occurred during the Late Pleistocene (MIS4- 2).