Richard C. Hulbert

A NEW MIDDLE EOCENE PROTOCETID WHALE (MAMMALIA: CETACEA: ARCHAEOCETI) AND ASSOCIATED BIOTA FROM GEORGIA

A shallow-marine fossil biota was recovered from the Blue Bluff unit (formerly part of the McBean Formation) in the Upper Coastal Plain of eastern Georgia. Biochronologically significant mollusks (e.g., Turritella nasuta, Cubitostrea sellaeformis, Pteropsella lapidosa) and calcareous nannoplankton (e.g., Chiasmolithus solitus, Reticulofenestra umbilica, Cribocentrum reticulatum) indicate a latest Lutetian-earliest Bartonian age, or about 40 to 41 Ma. Georgiacetus vogtlensis new genus and species is described from a well-preserved, partial skeleton. Georgiacetus is the oldest known whale with a true pterygoid sinus fossa in its basicranium and a pelvis that did not articulate directly with the sacral vertebrae, two features whose acquisitions were important steps toward adaptation to a fully marine existence. The posterior four cheek teeth of G. vogtlensis form a series of carnassial-like shearing blades. These teeth also bear small, blunt accessory cusps, which are regarded as being homologous with the larger, sharper accessory cusps

Revised age of the late Neogene terror bird (Titanis) in North America during the Great American Interchange

The giant flightless terror bird Titanis walleri is known from Florida and Texas during the late Neogene. The age of T. walleri is problematic because this taxon co-occurs with temporally mixed (i.e., time-averaged) faunas at two key sites. Thus, prior to this study, T. walleri from the Santa Fe River, Florida (type locality), was either as old as late Pliocene (ca. 2.2 Ma) or as young as latest Pleistocene (ca. 15 ka). Likewise, T. walleri from the Nueces River, Texas, was either early Pliocene (ca. 5 Ma) or latest Pleistocene (ca. 15 ka). In order to better resolve this age range, the rare earth element (REE) patterns of T. walleri from the Santa Fe River, Florida, were compared to two biochronologically distinctive groups (late Pliocene versus late Pleistocene) of fossil mammals from the same locality. Similarly, the REE patterns of T. walleri from Texas were compared to two groups (early Pliocene versus latest Pleistocene) of fossil mammals from the same locality. The REE patterns of T. walleri from Florida are indistinguishable from those of the co-occurring late Pliocene mammals. Likewise, the REE pattern of T. walleri from Texas is indistinguishable from those of the co-occurring early Pliocene mammals. Given these REE constraints, the revised age of T. walleri is early Pliocene in Texas (ca. 5 Ma) and late Pliocene (ca. 2.2–1.8 Ma) in Florida. As such, T. walleri is interpreted as an early immigrant during the Great American Interchange prior to the formation of the Isthmian land bridge. No evidence currently exists for Pleistocene T. walleri in North America.

THE STATUS OF THE DINOSAURIAN GENUS DICLONIUS AND THE TAXONOMIC UTILITY OF HADROSAURIAN TEETH

E. H. Sellards (1916, p. 96) applied the new binomen “ Hipparion minor ” to the very small hipparionine horse from the early Pliocene Bone Valley District of peninsular Florida. Since the gender of Hipparion is neuter, the ending of the species name must be amended to agree with it (Articles 32d and 34b, ICZN, 1985), and thus the correct spelling of the original species name for this taxon is Hipparion minus Sellards. This correction was previously instituted by Hay (1930, p. 693). However, this name is a primary junior homonym of Hipparion minus Pavlow, 1890, a Eurasian species of horse, and as such is permanently invalid (Articles 52b, 53c(ii), and 57b, ICZN, 1985). Hipparion minus Pavlow was in common use for small hipparionine horses from Russia and Greece during the first quarter of this century. Although now generally regarded as a nomen dubium, it was used as a valid subspecies as recently as Pirlot (1956). Its present disuse in no way affects the condition of primary homonymy between the two species names.

Giant Short-faced Bears (Arctodus simus) in Pleistocene Florida USA, a Substantial Range Extension

Abstract Fossils of the giant short-faced bear, Arctodus simus (Cope, 1879), have been recovered from over 100 localities in North America, extending from Mexico to Alaska and California to Virginia. Despite this large range, the species has never been recorded from the southeastern United States. The lesser short-faced bear, Arctodus pristinus Leidy, 1854 is well represented from this region, particularly Florida, but all known occurrences are late Pliocene – middle Pleistocene in age (about 2.5 to 0.3 Ma). Differentiating A. simus from A. pristinus can be difficult because large individuals of A. pristinus overlap in size with small individuals of A. simus, and there are few morphological differences. However, these two taxa can be clearly separated based on the relative proportions of their molars and premolars. Two Pleistocene records of A. simus representing a minimum of three individuals from the Withlacoochee River drainage of central Florida are reported here, substantially extending the distribution of this massive bear into southeastern North America. A late Pleistocene age for these occurrences is corroborated by an associated Rancholabrean fauna and rare earth elemental analyses. One of the reported individuals is quite large, supporting the hypothesis of extreme sexual dimorphism in A. simus and rejecting a hypothesis of two subspecies.

A New Machairodont from the Palmetto Fauna (Early Pliocene) of Florida, with Comments on the Origin of the Smilodontini (Mammalia, Carnivora, Felidae)

South-central Florida’s latest Hemphillian Palmetto Fauna includes two machairodontine felids, the lion-sized Machairodus coloradensis and a smaller, jaguar-sized species, initially referred to Megantereon hesperus based on a single, relatively incomplete mandible. This made the latter the oldest record of Megantereon, suggesting a New World origin of the genus. Subsequent workers variously accepted or rejected this identification and biogeographic scenario. Fortunately, new material, which preserves previously unknown characters, is now known for the smaller taxon. The most parsimonious results of a phylogenetic analysis using 37 cranio-mandibular characters from 13 taxa place it in the Smilodontini, like the original study; however, as the sister-taxon to Megantereon and Smilodon. Accordingly, we formally describe Rhizosmilodon fiteae gen. et sp. nov. Rhizosmilodon, Megantereon, and Smilodon ( =  Smilodontini) share synapomorphies relative to their sister-taxon Machairodontini: serrations smaller and restricted to canines; offset of P3 with P4 and p4 with m1; complete verticalization of mandibular symphysis; m1 shortened and robust with widest point anterior to notch; and extreme posterior “lean” to p3/p4. Rhizosmilodon has small anterior and posterior accessory cusps on p4, a relatively large lower canine, and small, non-procumbent lower incisors; all more primitive states than in Megantereon and Smilodon. The former also differs from Megantereon and Smilodon gracilis by having a very small mandibular flange. Rhizosmilodon is the oldest known member of the Smilodontini, suggesting that the tribe originated in North America. Two more derived, similar-sized species evolved in parallel during the Blancan, Megantereon hesperus and Smilodon gracilis. The former is rarer, known only from the north-central and northwestern US, and presumably dispersed into the Old World. The latter is known from the eastern and southern US, and dispersed into South America.