Rolf L. Aalbu

A REVISION OF THE ELEODES (SUBGENUS CAVERNELEODES) WITH NEW SPECIES AND NOTES ON CAVE BREEDING ELEODES (TENEBRIONIDAE: AMPHIDORINI)

Abstract. Subgenus Caverneleodes of the genus Eleodes is diagnosed and revised. Six new species from the United States: California (E. microps); Utah and Northern Arizona (E. wynnei), Central Arizona (E. wheeleri), Southern New Mexico (E. guadalupensis), and Mexico (E. thomasi and E. grutus) are described. The biogeography of the subgenus is discussed. Diagnoses and a key are provided to known species of Caverneleodes. Relationships with other Eleodes are discussed. Cave associated Amphidorini are surveyed.

A Revision of the Genus Eschatomoxys Blaisdell (Tenebrionidae: Pimeliinae: Edrotini) with Notes on the Biology

Abstract The tenebrionid genus Eschatomoxys Blaisdell is revised with descriptions of four new species, E. pholeter, E. paco, E. rosei, and E. andrewsi. Eschatomoxys wagneri Blaisdell, the type species, and E. tanneri Sorenson and Stones, are redescribed with illustrations and a key for the identification of all species. The biology of these rare beetles is discussed with emphasis on the cave dwelling species. The species have metapopulations fragmented among cave and island habitats.

The Tenebrionidae of California: A Time Sensitive Snapshot Assessment

Abstract Due to a diversity of habitats and its geologic history, the US state of California hosts a spectacular assemblage of darkling beetle species (Coleoptera: Tenebrionidae). In addition to being part of the California Floristic Province, one of 34 global biodiversity hotspots identified by Conservation International, California also has additional areas which are parts of the Great Basin, Mojave, and Sonoran deserts. California is divided into nine floristic regions. Each region is assessed in terms of faunal composition and endemism. A “snapshot” of our present knowledge of the Tenebrionidae indicates that 447 currently recognized species, representing 108 genera, occur in California of which one hundred and ninety are endemic. California is compared to other nearby regions in diversity and endemism. An analysis of currently valid species vs a more realistic species account based on unpublished records of likely synonyms and known species yet to be described in the scientific literature is presented. The California Floristic Region, rather than other more arid parts of California, has the highest number of total and endemic species. Because of their high diversity and endemism, tenebrionids could potentially provide a valuable tool for monitoring the environment for conservation purposes.

Catalogue of Tenebrionidae (Coleoptera) of North America

Abstract This catalogue includes all valid family-group (8 subfamilies, 52 tribes, 14 subtribes), genus-group (349 genera, 86 subgenera), and species-group names (2825 species, 215 subspecies) of darkling beetles (Coleoptera: Tenebrionidae) known to occur in North America1 and their available synonyms. Data on extant, subfossil and fossil taxa are given. For each name the author and year and page number of the description are provided, with additional information (e.g., type species for genus-group names, author of synonymies for invalid taxa) depending on the taxon rank. Several new nomenclatural acts are included. One new genus, Lepidocnemeplatia Bousquet and Bouchard, is described. Spelaebiosis Bousquet and Bouchard [for Ardoinia Özdikmen, 2004], Blapstinus marcuzzii Aalbu [for Blapstinus kulzeri Marcuzzi, 1977], and Hymenorus campbelli Bouchard [for Hymenorus oculatus Doyen and Poinar, 1994] are proposed as new replacement names. Supporting evidence is provided for the conservation of usage of Tarpela micans (Fabricius, 1798) nomen protectum over Tarpela vittata (Olivier, 1793) nomen oblitum. The generic names Psilomera Motschulsky, 1870 [= Stenomorpha Solier, 1836], Steneleodes Blaisdell, 1909 [= Xysta Eschscholtz, 1829], Ooconibius Casey, 1895 and Euconibius Casey, 1895 [= Conibius LeConte, 1851] are new synonyms (valid names in square brackets). The following 127 new synonymies of species-group names, listed in their original combination, are proposed (valid names, in their current combination, placed in square brackets): Bothrasida mucorea Wilke, 1922 [= Pelecyphorus guanajuatensis (Champion, 1884)]; Parasida zacualpanicola Wilke, 1922 [= Pelecyphorus asidoides Solier, 1836]; Stenosides kulzeri Pallister, 1954, Stenosides bisinuatus Pallister, 1954, and Parasida trisinuata Pallister, 1954 [= Pelecyphorus dispar (Champion, 1892)]; Asida favosa Champion, 1884 and Asida similata Champion, 1884 [= Pelecyphorus fallax (Champion, 1884)]; Ologlyptus bicarinatus Champion, 1884 [= Pelecyphorus indutus (Champion, 1884)]; Parasida laciniata Casey, 1912 and Parasida cristata Pallister, 1954 [= Pelecyphorus liratus (LeConte, 1854)]; Parasida esperanzae Wilke, 1922 and Parasida mixtecae Wilke, 1922 [= Pelecyphorus longipennis (Champion, 1884)]; Parasida tolucana Casey, 1912 [= Pelecyphorus scutellaris (Champion, 1884)]; Parasida purpusi Wilke, 1922 [= Pelecyphorus tristis (Champion, 1884)]; Astrotus nosodermoides Champion, 1892 [= Pelecyphorus erosus (Champion, 1892)]; Astrotus seticornis var. humeralis Champion, 1884 [= Pelecyphorus seticornis (Champion, 1884)]; Pactostoma breviuscula Casey, 1912, Pactostoma exoleta Casey, 1912, Pactostoma luteotecta Casey, 1912, Pactostoma monticola Casey, 1912, Pactostoma obtecta Casey, 1912, and Pactostoma sigillata Casey, 1912 [=Pelecyphorus anastomosis (Say, 1824)]; Ologlyptus canus Champion, 1884 and Ologlyptus sinuaticollis Champion, 1884 [= Pelecyphorus graciliformis (Solier, 1836)]; Gonasida elata reducta Casey, 1912, Gonasida elata prolixa Casey, 1912, and Gonasida aucta Casey, 1912 [= Philolithus elatus compar (Casey, 1912)]; Gonasida alaticollis Casey, 1912 [= Philolithus elatus difformis (LeConte, 1854)]; Gonasida gravida Casey, 1912 [= Philolithus elatus elatus (LeConte, 1853)]; Pelecyphorus aegrotus limbatus Casey, 1912 [= Philolithus aegrotus aegrotus (LeConte, 1861)]; Pelecyphorus corporalis Casey, 1912, Pelecyphorus reptans Casey, 1912, Pelecyphorus socer Casey, 1912, Pelecyphorus abscissus Casey, 1912, Pelecyphorus fumosus Casey, 1912, Pelecyphorus parvus Casey, 1912, Pelecyphorus morbillosus pacatus Casey, 1912, Pelecyphorus morbillosus sobrius Casey, 1912, Pelecyphorus piceus Casey, 1912, Pelecyphorus piceus crudelis Casey, 1912, Pelecyphorus snowi Casey, 1912, and Pelecyphorus subtenuis Casey, 1912 [= Philolithus morbillosus (LeConte, 1858)]; Bothrasida sanctae-agnae Wilke, 1922 [= Stenomorpha funesta (Champion, 1884)]; Asida flaccida Horn, 1896 [= Stenomorpha embaphionides (Horn, 1894)]; Asida angustula Casey, 1890, Stethasida stricta Casey, 1912, Stethasida muricatula languida Casey, 1912, Stethasida pertinax Casey, 1912, Stethasida socors Casey, 1912, Stethasida angustula inepta Casey, 1912, Stethasida tenax Casey, 1912, and Stethasida vegrandis Casey, 1912 [= Stenomorpha muricatula (LeConte, 1851)]; Stethasida obsoleta expansa Casey, 1912, Stethasida obsoleta opacella Casey, 1912, Stethasida brevipes Casey, 1912, Stethasida torpida Casey, 1912, Stethasida convergens Casey, 1912, Stethasida discreta Casey, 1912, Stethasida longula Casey, 1912, Stethasida adumbrata Casey, 1912, Stethasida occulta Casey, 1912, Stethasida tarsalis Casey, 1912, Stethasida unica Casey, 1912, and Pelecyphorus laevigatus Papp, 1961 [= Stenomorpha obsoleta (LeConte, 1851)]; Trichiasida eremica Wilke, 1922 [= Stenomorpha difficilis (Champion, 1884)]; Trichiasida lineatopilosa Casey, 1912 [= Stenomorpha hirsuta (LeConte, 1851)]; Trichiasida tenella Casey, 1912 [= Stenomorpha hispidula (LeConte, 1851)]; Trichiasida duplex Casey, 1912 [= Stenomorpha villosa (Champion, 1884)]; Alaudes squamosa Blaisdell, 1919, Alaudes testacea Blaisdell, 1919, and Alaudes fallax Fall, 1928 [= Alaudes singularis Horn, 1870]; Edrotes barrowsi Dajoz, 1999 [=Edrotes ventricosus LeConte, 1851]; Nyctoporis tetrica Casey, 1907 and Nyctoporis maura Casey, 1907 [= Nyctoporis aequicollis Eschscholtz, 1831]; Nyctoporis pullata Casey, 1907 [= Nyctoporis sponsa Casey, 1907]; Eleodes tibialis forma oblonga Blaisdell, 1909 [= Eleodes tibialis Blaisdell, 1909]; Eleodes (manni var.) variolosa Blaisdell, 1917 [= Eleodes constrictus LeConte, 1858]; Eleodes cordata forma sublaevis Blaisdell, 1909, Eleodes cordata forma intermedia Blaisdell, 1909, Eleodes cordata forma oblonga Blaisdell, 1909, Eleodes cordata forma elongata Blaisdell, 1909, and Eleodes (cordata var.) adulterina Blaisdell, 1917 [= Eleodes cordata Eschscholtz, 1829]; Eleodes hornii var. monticula Blaisdell, 1918 and Eleodes manni sierra Blaisdell, 1925 [= Eleodes fuchsii Blaisdell, 1909]; Eleodes parvicollis var. squalida Blaisdell, 1918 [= Eleodes parvicollis Eschscholtz, 1829]; Eleodes reflexicollis Mannerheim, 1843 and Eleodes parvicollis forma farallonica Blaisdell, 1909 [= Eleodes planata Eschscholtz, 1829]; Eleodes indentata Blaisdell, 1935 [= Eleodes rotundipennis LeConte, 1857]; Eleodes intricata Mannerheim, 1843 [= Eleodes scabrosa Eschscholtz, 1829]; Eleodes horni fenyesi Blaisdell, 1925 [= Eleodes tenebrosa Horn, 1870]; Eleodes cordata var. horrida Blaisdell, 1918 [= Eleodes tuberculata Eschscholtz, 1829]; Eleodes oblonga Blaisdell, 1933 [= Eleodes versatilis Blaisdell, 1921]; Eleodes dentipes marinae Blaisdell, 1921 [= Eleodes dentipes Eschscholtz, 1829]; Eleodes carbonaria forma glabra Blaisdell, 1909 [= Eleodes carbonaria carbonaria (Say, 1824)]; Eleodes granosa forma fortis Blaisdell, 1909 [= Eleodes granosa LeConte, 1866]; Eleodes pilosa forma ordinata Blaisdell, 1909 [= Eleodes pilosa Horn, 1870]; Trogloderus costatus pappi Kulzer, 1960 [= Trogloderus tuberculatus Blaisdell, 1909]; Trogloderus costatus mayhewi Papp, 1961 [= Trogloderus vandykei La Rivers, 1946]; Bolitophagus cristatus Gosse, 1840 [= Bolitotherus cornutus (Fabricius, 1801)]; Eleates explanatus Casey, 1890 [= Eleates depressus (Randall, 1838)]; Blapstinus sonorae Casey, 1890 [= Blapstinus brevicollis LeConte, 1851]; Blapstinus falli Blaisdell, 1929 [= Blapstinus castaneus Casey, 1890]; Blapstinus brunneus Casey, 1890 and Blapstinus coronadensis Blaisdell, 1892 [=Blapstinus histricus Casey, 1890]; Blapstinus hesperius Casey, 1890 [=Blapstinus intermixtus Casey, 1890]; Blapstinus cinerascens Fall, 1929 [= Blapstinus lecontei Mulsant and Rey, 1859]; Blapstinus niger Casey, 1890 and Blapstinus cribricollis Casey, 1890 [= Blapstinus pimalis Casey, 1885]; Blapstinus arenarius Casey, 1890 [= Blapstinus pratensis LeConte, 1859]; Blapstinus gregalis Casey, 1890 [= Blapstinus substriatus Champion, 1885]; Blapstinus hydropicus Casey, 1890 [= Blapstinus sulcatus LeConte, 1851]; Blapstinus hospes Casey, 1890 [= Blapstinus vestitus LeConte, 1859]; Notibius reflexus Horn, 1894 [= Conibius opacus (LeConte, 1866)]; Notibius affinis Champion, 1885 [=Conibius rugipes (Champion, 1885)]; Conibius parallelus LeConte, 1851 [= Conibius seriatus LeConte, 1851]; Nocibiotes rubripes Casey, 1895 [=Nocibiotes caudatus Casey, 1895]; Nocibiotes gracilis Casey, 1895 and Nocibiotes acutus Casey, 1895 [=Nocibiotes granulatus (LeConte, 1851)]; Conibius alternatus Casey, 1890 [= Tonibius sulcatus (LeConte, 1851)]; Pedinus suturalis Say, 1824 [= Alaetrinus minimus (Palisot de Beauvois, 1817)]; Menedrio longipennis Motschulsky, 1872 [= Tenebrio obscurus Fabricius, 1792]; Hymenophorus megops Hatch, 1965 and Telesicles magnus Hatch, 1965 [= Hymenorus sinuatus Fall, 1931]; Andrimus concolor Casey, 1891 and Andrimus convergens Casey, 1891 [= Andrimus murrayi (LeConte, 1866)]; Mycetochara marshalli Campbell, 1978 [= Mycetochara perplexata Marshall, 1970]; Phaleria globosa LeConte, 1857 [= Phaleria picta Mannerheim, 1843]. The following subspecies of Trogloderus costatus LeConte, 1879 are given species rank: Trogloderus nevadus La Rivers, 1943, Trogloderus tuberculatus Blaisdell, 1909, and Trogloderus vandykei La Rivers, 1946. The following taxa, previously thought to be junior synonyms, are considered valid: Amphidora Eschscholtz, 1829; Xysta Eschscholtz, 1829; Helops confluens (Casey, 1924). Two new combinations are proposed: Stenomorpha spinimana (Champion, 1892) and Stenomorpha tenebrosa (Champion, 1892) [from the genus Parasida Casey, 1912]. The type species [placed in square brackets] of the following 12 genus-group taxa are designated for the first time: Lagriola Kirsch, 1874 [Lagriola operosa Kirsch, 1874]; Locrodes Casey, 1907 [Emmenastus piceus Casey, 1890]; Falacer Laporte, 1840 [Acanthopus cupreus Laporte, 1840 (= Helops contractus Palisot de Beauvois, 1812)]; Blapylis Horn, 1870 [Eleodes cordata Eschscholtz

Reinstatement of Eschatoporiini Blaisdell, 1906, a unique tribe of blind cavernicolous Tenebrionidae from California, with a new species from Napa County (Coleoptera, Tenebrionidae, Lagriinae)

Abstract The tribe Eschatoporini Blaisdell, 1906 is reinstated, based on molecular and morphological data, and the spelling corrected as Eschatoporiini. The tribe currently includes only the cave-dwelling genus Eschatoporis Blaisdell, 1906 from California, which is associated with underground aquifers. A second species of Eschatoporis is described from a cave in Napa County, California. The phylogenetic placement of Eschatoporiini within the Lagriinae is examined, and notes on the biology of Eschatoporis are provided.

Opatroides punctulatus Brullé now established in California (Coleoptera: Tenebrionidae).

Abstract Opatroides punctulatus Brullé, 1832 (Coleoptera; Tenebrionidae; Opatrini), is reported established for the first time in the New World in California near Sacramento. A key to distinguish Opatroides from species of similar genera is provided, with images and a brief diagnosis of O. punctulatus. The potential spread and pest status of the beetle should be monitored.

Caribanosis gen. nov. from Hispaniola (Pimeliinae: Stenosiini) with taxonomic notes on the tribes belopini and stenosini (Coleoptera: Tenebrionidae)

Abstract. Rhypasma Pascoe, 1862 is transferred from the tribe Stenosini (Pimeliinae) to Belopini (Lagriinae). Caribanosis gen. nov. is described and placed in Stenosini (Tenebrionidae: Pimeliinae) to hold Rhypasma quisqueyanus Garrido et Varela, 2011. The following new combination is established: Caribanosis quisqueyanus (Garrido et Varela, 2011), comb. nov. Caribanosis is similar to the South American genus Grammicus Waterhouse, 1845 but differs in having a single central pronotal keel, not two lateral keels as in Grammicus. Both are members of the subtribe Stenosina.

A Revision of Craniotus Leconte (Coleoptera: Tenebrionidae: Pimeliinae: Asidini), with Descriptions of New Insular Species from Mexico and Notes on Distribution and Biology

Abstract The genus Craniotus LeConte is revised. The genus includes Craniotus pubescens LeConte and the following new species: Craniotus mardecortesi Aalbu, Smith, and Sanchez Piñero (Isla San Esteban, Baja California, Mexico) and Craniotus triplehorni Aalbu, Smith, and Sanchez Piñero (Gulf Islands and Baja California, Mexico). Craniotus blaisdelli Tanner is placed as a synonym of C. pubescens. This form corresponds to northern populations of C. pubescens. A key to the species is provided. The distribution and biology of the genus are briefly discussed.

Charles A. Triplehorn: An Inordinate Fondness for Darkling Beetles

Charles A., “Chuck to his friends”, Triplehorn was born at Bluffton, Ohio on 27 October 1927. Active in the Boy Scouts as a youth, he attained the rank of Eagle Scout. His interest in wildlife, especially reptiles, lead him to the life sciences in college. He graduated with a Bachelors degree in zoology and then a Masters degree in entomology (under thesis advisor Joseph Knull) from the Ohio State University in 1949 and 1952, respectively. His first paying job in entomology was at the Delaware Agricultural Experiment Station, where he worked on the insects of oak trees. After a two year stint, he returned to schooling, earning a PhD in entomology at Cornell University under Dr. Howard Evans in 1957, with a dissertation on the systematics of darkling beetles. In 1962, he moved back to his home state to take a position with the Ohio Agricultural Experiment Station, where he worked on corn insects. During 1964–1966, Chuck was stationed in Brazil with US-AID. While the work on corn insects displaced his interest in snakes, the former was eventually displaced by his interest in darkling beetles, a field in which he became the world’s foremost authority. Dedicated to students, to his colleagues, and his profession, Chuck was elected President of The Coleopterists Society in 1976 and later President of the Entomological Society of America in 1985. Among his honors are his election to honorary membership of both The Coleopterists Society and the Entomological Society of America and the latter society’s highest honor – the Founders Memorial Award. As a faculty member at the Ohio State University, Chuck was asked by his former mentors Donald Borror and Dwight DeLong to co-author the fourth edition of their classic textbook, An Introduction to the Study of Insects. It was not Chuck’s first involvement in the book. As a graduate student, he had contributed his own illustration of Merinus laevis to the first edition. From his initial involvement as editor of the Coleoptera chapter, he eventually became the chief “compiler” for the modern editions. It has become the standard textbook for entomology throughout the world, with editions in mutiple languages including Arabic and Indonesian. It was this textbook, perhaps more than any of his many publications, that brought him fame. As for fortune, there were, of course, the royalties, which Chuck once described as “I could have made more per hour bagging groceries or pumping gas.”

Editorial: Third International Tenebrionoidea Symposium

The Third International Tenebrionoidea Symposium (ITS) was held at Arizona State University in Tempe, Arizona USA on August 7th and 8th, 2013. Researchers from ten countries participated with a total of 36 attendees (Figure 1). It was the first formal meeting of the international tenebrionoid research community since the October 2005 in Lyon, France. Though the previous meetings did not list themselves as the beginning of a series, we consider it fitting to acknowledge them as the first two modern international meetings specifically organized for the sharing and dissemination of Tenebrionoidea research. Figure 1. August 7th, before the first talk. The 1st International Tenebrionid Symposium, entitled “Systematics and Biogeography of Tenebrionoidea”, was held in 2002 at the Zoologisches Staatssammlung, Munchen (Germany) to honor Dr. Hans J. Bremer’s work on tenebrionids and celebrate the museum’s acquisition of his collection. This event organized by Dr. Martin Baehr resulted in a highly successful meeting. The 2nd International Tenebrionoidea Symposium, entitled “Coleoptera Tenebrionoidea: Taxonomy, Biogeography, and Faunistics”, was held in 2005 at the Lyon Museum (France) following the acquisition of the remarkable tenebrionid collection of Jaroslav Picka. Following the symposium, many of the presentations were published in Cahiers Scientifiques (Fascicule 10). Again a highly successful meeting this time organized by Dr. Harold LaBrique. To continue this successful tradition, and encourage tenebrionoid workers from around the world to meet, share their research, and form new collaborations, researchers in the US and Canada decided to host the 3nd International Tenebrionoidea Symposium. A steering committee was assembled with representatives from Arizona State University, California Academy of Sciences, the Canadian National Collection of Insects, and the Smithsonian Institution. Arizona State University in Tempe, Arizona was ultimately chosen to host the symposium due to its institutional support, excellent facilities, and multiple opportunities for field work both before and after the meeting. Presentations were given on August 7th and 8th, 2013. Before the meeting, researchers visited US collections on both the west and east coasts and held a pre-meeting collecting trip through California, Nevada, Utah, and Arizona. Gustavo Flores had the most impressive itinerary of museum visits. After flying into New York City from Mendoza, Argentina, Gustavo visited the American Museum of Natural History (AMNH – New York, New York), the Smithsonian Institution (NMNH – Washington, D.C.), the C.A. Triplehorn Insect Collection at Ohio State University (OSUC – Columbus, Ohio), the Field Museum (FMNH – Chicago, Illinois), and Rolf Aalbu’s personal collection (RLAC – El Dorado Hills, California). In Sacramento, Gustavo joined Wolfgang Schawaller, Roland Grimm, and Rene Fouque who had been working in the California Academy of Sciences (CASC – San Francisco, California), California Department of Food and Agriculture (CDFA – Sacramento, California), and RLAC collections the prior week. Rolf, Gustavo, Rene, Roland, and Wolfgang then drove from Sacramento to Tempe while doing field work through California, Nevada, Utah, and north central Arizona (Figure 2). Figure 2. Pre-meeting sightseeing stop at Zion National Park, Utah. Left to right: Wolfgang Schawaller, Rene Fouque, Gustavo Flores. During the meeting 21 presentations, seventeen 20-minute talks and four posters, where given (see http://insectbiodiversitylab.org/3ITS_presentations.html) ranging from species-level revisions to broad scale Tenebrionidae phylogenies and inventories, darkling beetles intercepted by USDA-APHIS during agricultural quarantine interceptions, and the first steps towards the construction of a Coleopteran Anatomy Ontology. Presentations were generally well received and elicited animated question and answer sessions. Many of the attendees had previously corresponded by email, but never met in person. For example, Guodong Ren’s research group (Figure 3) has been remarkably productive, but this was the first time any of the American (North and South) or European visitors were able to meet him face to face. Others, such as Chuck Triplehorn (Figure 4) are well known to almost all attendees through both research and previous visits. Following the first day’s presentations, Bill Warner led an evening collecting expedition to Oak Flat Campground in Pinal County. Figure 3. Visiting Chinese and US-based Chinese researchers. Left to right: Yuxia Yang, Li Zhong, Guodong Ren, Guanyang Zhang (ASU postdoc), Shanshan Liu, Caixia Yuan Figure 4. Dr. Charles A. Triplehorn showing off a Triplehornia metallica Matthews and Lawrence shirt made by his grandson. Group discussions were also held during the symposium on potential large scale tenebrionid projects that could be undertaken as a community, the organization of a Proceedings volume from the Symposium, collecting localities for the post-meeting trip, and potential localities and dates for the Fourth International Tenebrionoidea Symposium. Informal talks on these and other tenebrionoid related matters extend far into the evening and past the closing session on August 8th (Figure 5). Pat Bouchard agreed to act as lead editor for a Proceedings volume in the journal Zookeys, for which we were and remain very grateful. Most articles included in this resulting special issue of ZooKeys are based on the contents of presentations during the Third International Tenebrionoidea Symposium, although papers submitted by all attendants were also welcome. Figure 5. Post meeting dinner. Left to right, back row: Bill Warner, Rich Cunningham, Pat Bouchard, Wolfgang Schawaller, Aaron Smith, Milton Campbell, Andrew Johnston, Marcin Kaminski; front row: Ron Somerby, Gael Kergoat, Roland Grimm, Rolf Aalbu, Rebecca ... After the formal symposium, attendees went their separate ways, with some doing solo collecting and some visiting US museums (California Academy of Sciences and the University of Arizona Insect Collection to name just two). Twelve researchers from five countries went to the Beetle Infestation VI on August 10th hosted by Pat and Lisa Sullivan in Ramsey Canyon, Huachuca Mountains, one of the most biologically diverse localities in the United States, before collecting through southern and central Arizona (Figures 6 & 7) eventually disbursing into smaller field groups or heading home. While a full tally of tenebrionoid species collected in association with the symposium will likely never be assembled, the first author collected approximately 40 darkling beetles species during and after the meeting. Most of the species collected can be sight IDed, at least to genus, using Bugguide. Figure 6. Post meeting collecting. Marcin Kaminski and Andrew Johnston near Madera Canyon. Figure 7. Post meeting afternoon break at Fred Skillman’s house, Cochise, AZ. Left to right: Wolfgang Schawaller, Pat Bouchard, Rolf Aalbu, Kojun Kanda, Andrew Johnston, Fred Skillman, Warren Steiner, Marcin Kaminski, Rene Fouque. ... Many of the presentations, a list of collecting localities, and additional pictures from the symposium are online at: http://www.insectbiodiversitylab.org/3ITS.html. Two researchers graciously volunteered to host the next symposium at their institutions: Gustavo Flores (CCT CONICET – Mendoza, Argentina) for 2016, or Guodong Ren (Hebei University – Baoding City, China) for 2015, and presented short talks highlighting the advantages of their respective cities. A survey was set up to allow the attendees of the Third symposium, current tenebrionoid researchers (those with at least one tenebrionioid manuscript in print), and graduate students working on tenebrionoids to vote for the host city of the Fourth International Tenebrionoidea Symposium. Voting was open until September 30th, 2013 and turnout was excellent. After over a month of voting, Mendoza, Argentina was chosen to host the next meeting in November 2015. See you in Mendoza!