Ronald A. Pursell

A Revision of the Infrageneric Taxa of Fissidens

Abstract The Fissidentaceae are recognized to include a single genus, Fissidens, which is divided into four subgenera, Aloma, Fissidens, Octodiceras, and Pachyfissidens, on the basis of new taxonomically useful characters i.e., peristome type, costa type, and number of files of exothecial cells. Subgenera Aloma and Octodiceras are not subdivided. Subgenus Fissidens consists of sections Fissidens and Sarawakia, comb. nov. Subgenus Pachyfissidens is divided into three sections: Amblyothallia, comb. nov., Crispidium comb. nov., and Pachyfissidens. The most primitive species of the genus are found in section Amblyothallia, while the most advanced species are in subgenera Aloma and Octodiceras. A key to the subgenera and sections is provided.

Taxonomic Notes on Neotropical Fissidens

One hundred and thirty-three names of Fissidens Hedw. (including Conomitrium Mont., Moenkemeyera C. Miill., and Simplicidens Herz.) are placed in the synonymy of 35 other names of Fissidens. Seventy-nine names are lectotypified. Neotypes are designated for F. amoenus C. Miill., F. distichellus Besch., F. oerstedianus C. Miill., F. pellucidus Hornsch., and F. serratus

A Conspectus of the Mosses of Antarctica, South Georgia, the Falkland Islands and Southern South America

A SURVEY is presGnted of thG taxa of mossGS which havG baan raportad from Antarctica, tha islands of tha Scotia Ridga, South Gaorgia, tha Falkland Islands and southam South Amarica, i.a. Argantina, Chila and Uruguay, including Juan Famandaz, from aarliast timas up to tha and of 1977. Tha rasults show that 1727 spacias and 254subspacias or variatias ara currantly accaptad for tha araa; of thasa 254 spacias and 40 subspecies or variatias are baliavad to be andamic. Nomanclatura is based on that in index muscorum (van dar Wijk and others, 1959,1962,1964, 1967,1969) or in later treatments where appropriate but in four cases, viz. Racomitrium crispulum, tha Polytrichacaaa, Bryum antarcticum and Sphagnum, reasons are given for adopting a different 3pproach. Tha list of synonyms includes tha basionym and those epithets which have baan used for material from tha araa but is not nomanclaturally exhaustive. Some 60 nominanuda ara also listed since specimens bearing thasa names ara frequently encountered in herbaria. Distribution follows tha worldwide schema adopted in Index muscorum grouped hare under six headings, viz. Juan Fernandez, southam South Amarica, tha Falkland Islands, South Gaorgia, Antarctic Peninsula region and Antarctic continental region, tha records for tha South Orkney Islands being included with tha peninsula region. It was found that 41 of tha taxa listed in Index muscorum as occurring in tha araa could not be substantiated from any other source and so have baan delated from tha list. As genera and species ara revised, further reductions in the totals of currently accepted taxa can be expected, although further field work may add others. The list of references is extensive and includes about 900 entries, all of which have been examined by the author. Included in the list is the literature relating to the basionyms. Where appropriate, both bibliographic and effective dates have been given to facilitate literature retrieval and taxonomic requirements.

Recent literature on bryophytes—110(3)

] Werner, J. 2006. Observations bryologiques au Luxembourg (20 serie): Especes remarquables observees en 2005 et revisions partielles recentes. Bulletin de la Societe des Naturalistes Luxembourgeois 107: 27–30. Werner, O., R. M. Ros & B. Goffinet. 2007. A reconsideration of the systematic position of Goniomitrium (Funariaceae) based on chloroplast sequence markers. The Bryologist 110: 108– 114. [New: Clavitheca nom. nov., C. poeltii (Ochyra) comb. nov.] Wigginton, M. J. 2006. Bryophytes of St Helena, South Atlantic Ocean. 1. Three new species of Cololejeunea (Jungermanniales, Lejeuneaceae), C. dianae sp. nov., C. sanctae-helenae sp. nov. and C. grossestyla sp. nov. Journal of Bryology 28: 363–373. Wilbraham, J. 2007. [Abstract] Mosses and the African Plants Initiative: digitisation at the Natural History Museum, London. Field Bryology 91: 39. Wilson, R., S. R. Gradstein, H. Schneider & J. Heinrichs. 2007. Unravelling the phylogeny of Lejeuneaceae (Jungermanniopsida): evidence for four main lineages. Molecular Phylogenetics and Evolution 43: 270–282. [Two subfamilies recognized, Ptychanthoideae and Lejeuneoideae; in the latter 3 clades are recognized, Lejeuneeae, Brachiolejeuneeae and Symbiezidium.] Wright, J. A. 2007. Reports of local meetings: Southern Group. Field Bryology 91: 60–61. Wu, Yu-Huan & Chien Gao. 2007. Apomarsupella verrucosa (Nichols.) Vaňa and its new synonym. Bulletin of Botanical Research 27: 3–5. [In Chinese with English abstract; Gymnomitrion papillosum is a new synonym.] Wynns, J. 2006. Taxonomic Studies in the Aquatic Moss Genus Platyhypnidium (Brachytheciaceae). xiii þ 224 pp. Appalachian State University, Boone, NC. [Master’s thesis; Platyhypnidium is polyphyletic; P. pringlei belongs in Oxyrrhynchium, as does Eurhynchium selaginellifolium.] Xie, Chun-Feng, Jian-Bo Qu, Bin Sun, Huai-Fang Guo & HongXiang Lou. 2007. Dumhirone A, an unusual phenylethyl cyclohexadienone from the Chinese liverwort Dumortiera hirsuta. Biochemical Systematics and Ecology 35: 162–165. Xiong, Yuan-Xin, Cui-Zhen Wu, Xiao-Yu Wang & Xiao-Li Yan. 2006. Taxonomic and geographic study of genus Horikawaea Nog. and H. nitida Nog. as a new record from Guizhou. Buck et al.: Recent literature on bryophytes 575

The Moss Genus Fissidens in New Zealand. An Illustrated Key

J. BEEVER, B. MALCOLM AND N. MALCOLM. The Moss Genus Fissidens in New Zealand. An Illustrated Key. Te Puninga Fissidens I Aotearoa. iv 1 91 pages (soft cover, coil-bound) 1 CD (pdf format). Micro-Optics Press, Box 320, Nelson, New Zealand. 2002. [ISBN 0-9582224-6-0.] Price: NZ

Fissidens sublimbatus, New to Morocco and the Canary Islands

60. (Price includes postage to anywhere in the world. A personal or institutional cheque in the buyer’s local currency for the equivalent amount is acceptable. Cheques should be made payable to Micro-Optics Press.)

Typification of Hedwig's Species of Fissidens

Abstract Fissidens sublimbatus is reported new to Morocco and the Canary Islands (Spain). Differences with F. arnoldii, F. bambergeri, F. crispus, and F. megalotis subsp. helictocaulos are given. A map showing the world distribution of the species is included.

Taxonomic Notes on Neotropical Fissidens. II. An Addendum

Type specimens of ten species of Fissidens, viz., F. adianthoides, F. asplenioides, F. bryoides, F. exilis, F. osmundoides, F. palmatus, F. polypodioides, F. semicompletus, F. subba- silaris and F. taxifolius, described by Hedwig in Species Muscorum (1801) and currently retained within the genus are discussed. Specimens later added to the Hedwig-Schwaegrichen Herbarium by Schwaegrichen are reevaluated taxonomically. The importance of the works of Johannes Hedwig to the taxonomy of mosses was formally recognized in 1910 when the International Botanical Congress selected his posthumously published Species mus- corum frondosorum descriptae et tabulis aeneis LXXVII coloratis illustratae (1801), hereafter re- ferred to as Species Muscorum, to be the starting point for the nomenclature of mosses, except Sphagnum (see Sayre 1959; Florschiitz 1960; Mar- gadant 1968). This decision has made the typifi- cation of those species described in Species Mus- corum of the utmost practical importance. Typification of the Hedwigian species, however, has not always been an easy task (see Florschiitz 1960; Margadant 1968; Koponen 1979 for elaboration). Typification of these species naturally centers around the specimens in the Hedwig-Schwaegrichen Herbarium now maintained in the Conservatoire et Jardin botaniques de Geneve (G). In his introduction to the fascimile printing of Species Muscorum, Flor- schiitz (1960) presents a short history of this her- barium. Presently it consists of approximately 1,200 sheets. However, some specimens are known to have been distributed to other herbaria and some spec- imens have probably been lost (see Florschiitz 1960; Stafleu and Cowan 1976-1983; Koponen 1979). Al-

Collecting the Knot-Hole Moss (Anacamptodon splachnoides)

Twenty-five names in Fissidens Hedw. (including Conomitrium Mont.) and one name in Eustichia (Brid.) Brid. are newly placed in the synonymy of 11 accepted names of Fissidens. Four names, Conomitrium generalium C. Mill. [= Fissidens generalium (C. Miill.) Broth.], Diplostichum ulei C. Mill., [= Eustichia ulei (C. Mill.) Paris], Fissidens fossicola C. Miall., and Fissidens moritzianus C. Miill. are lectotypified here. A lengthy list of synonyms and other taxonomic changes on neotropical Fissidens was published recently in preparation of a monograph of the genus in the neotropics (Pursell 1994b). This report is an addendum to that list. Four lectotypes have been designated. FISSIDENS ANGUSTE-LIMBATUS Mitt., J. Linn. Soc., Bot. 12: 601. 1869, non Fissidens angusti-limbatus Broth., Hedwigia 38: 210. 1899. Type listed in Pursell (1994b). Fissidens fossicola C. Mill., Linnaea 9: 354. 1882, syn. nov. PROTOLOGUE: Argentinia Buenos-Airensis, in fossa Tigre, 28. Martio 1880: Prof. O. Schnyder. TYPE: ARGENTINA. Buenos-Airensis: in fossa Tigre, 28 Martio 1880, O. Schnyder 55 (lectotype, designated here BM). Fissidens anguste-limbatus is typically aquatic, although at times it can be found stranded well above the water line. Its aquatic habitat, and ovate to oblong-ovate, mucronate leaves will help to distinguish this species from others in section Fissidens. As is characteristic of species in section Fissidens, the limbidium is variable in thickness and

Fissidens jansenii (Fissidentaceae; Bryopsida), a New Aquatic Species from Portugal

Abstract. The “knot-hole moss” [Anacamptodon splachnoides (Fröhl. ex Brid.) Brid.] is seldom collected by bryologists, and is considered uncommon or even rare. However, 24 collections of A. splachnoides were made from trunks of red maple (Acer rubrum L.), sugar maple (Acer saccharumMarsh.), and chestnut oak (Quercus montana Willd.), as well as from the upper pilear surface of basidiomata (“conks”) of the “mossy-cap polypore” [Oxyporus populinus Fr. = Fomes connatus (Weinm.) Gillet]. Most collections were made in Pennsylvania. Of the 24 collections, 11 were from moist knotholes, seven from moist decayed hollows or crevices, three from limb/trunk crotches, and three from mossy-cap polypore conks. Twenty-three of the 24 collections were made during the summer of 2006. Anacamptodon splachnoides was found in both closed forests and open parks, and at both high and low elevations. These results indicate that A. splachnoides is not uncommon, but overlooked because of its specific habitat.