Shih Chungkun

New Species of Cimbrophlebiidae (Insecta: Mecoptera) from the Middle Jurassic of Northeastern China

Four new species of three genera in the family Cimbrophlebiidae, Telobittacus decorus sp. nov., Bellicimbrophlebia heteroneura sp. nov., Cimbrophlebia amoena sp. nov., and Cimbrophlebia gracilenta sp. nov., are described and illustrated based on eleven well-preserved fossil specimens. These specimens were collected from the latest Middle Jurassic Jiulongshan Formation of Daohugou, Inner Mongolia, China. This is the first record of the genus Cimbrophlebia in the Jurassic extending its age from the Lower Cretaceous to the Middle Jurassic. Due to same venational characters, we propose that an undescribed fossil specimen of Cimbrophlebiidae from the Jurassic of Germany is attributed to C. gracilenta sp. nov. Our new findings demonstrate an evolutionary trend of venational change from 185 to 50 Ma that the level of the bifurcation of Rs is gradually moving toward the base of wing in reference to the bifurcation of M.

A New Mesopsychid (Mecoptera) from the Middle Jurassic of Northeastern China

: The family Mesopsychidae Tillyard, 1917 presently consists of ten described genera from the Early Permian to the Early Cretaceous of Australia, China, Kyrgyzstan, Russia, South Africa, Tajikistan, and Ukraine. Herein, a new genus and a new species of fossil mesopsychid, Epicharmesopsyche pentavenulosa gen. et sp. nov., is described from the Middle Jurassic Jiulongshan Formation of Inner Mongolia, China, supplementing the family-level diversity. The well-preserved, new material reveals many characters, including antennae, wing venation, shape and genitalia that increase our morphological understanding of the family, although unfortunately the mouthparts and most of the legs are not preserved. Diagnosis of the Mesopsychidae is emended to include a four- or five-branched MP in both the fore- and hind wings. This is the first documentation of a wing coupling structure in the Mesopsychidae, consisting of three to four frenula bristles on the humeral lobe at the base of the costal margin of both hind wings. Asymmetrical shape and size of the left and right wings on a specimen of E. pentavenulosa gen. et sp. nov., seems to be a common condition for mesopsychid taxa from northeastern China.

A New Axymyiid Genus with Two New Species from the Middle Jurassic of China (Diptera: Nematocera: Axymyiidae)

: The Axymyiidae is one of the small families of the suborder Nematocera within the Diptera. Up to date, three genera and seven species of extant axymyiids have been described from the Holarctic Region, including Canada, China, Hungary, Japan, Russia, and USA, with three genera and three species of fossil taxa described from the Middle Jurassic of Daohugou, northeastern China. A new genus Raraxymyia gen. nov. and two new species, R. parallela and R. proxima gen. et sp. nov. from the Middle Jurassic Jiulongshan Formation of Daohugou Village, Inner Mongolia, China, are herein described based on their well-preserved wings and body characters. A revised key to the genera of axymyiids is provided.

New Species of Cretolimonia and Mesotipula(Diptera:Limoniidae) from the Middle Jurassic of Northeastern China

Two new species, Cretolimonia excelsa sp. nov. and Mesotipula gloriosa sp. nov., in Limoniidae of Diptera are described and illustrated, based on three fossil specimens with bodies and complete wings. These specimens were collected from the latest Middle Jurassic Jiulongshan Formation at Daohugou, Inner Mongolia, China. These are the first fossil records of these two genera from China.