Torsten Utescher

The coexistence approach -- a method for quantitative reconstructions of Tertiary terrestrial palaeoclimate data using plant fossils

Abstract The coexistence approach is introduced as a method for quantitative terrestrial palaeoclimate reconstructions in the Tertiary. It is based on the assumption that Tertiary plant taxa have similar climatic requirements to their nearest living relatives. The aim of the coexistence approach is to find for a given fossil flora and a given climate parameter the climatic interval in which all nearest living relatives of the fossil flora can coexist. For this purpose we have developed a data base, CLIMBOT (containing for over 800 Tertiary plant taxa, the nearest living relatives and their climatic requirements) and an algorithm for analysis, CLIMST. For all types of fossil floras the coexistence approach thus allows the rapid quantitative reconstruction of 10 different climate parameters, including the mean annual temperature, temperature of the warmest month, temperature of the coldest month, mean annual precipitation, maximum monthly precipitation, minimum monthly precipitation, precipitation of the warmest month, relative humidity, potential evaporation, as well as the ratio of mean annual precipitation over potential evaporation. The reliability and resolution of the coexistence approach are tested with various techniques and proved to be reasonably good; for instance, the resolution with respect to the mean annual temperature can be up to 1°C. Applications to various modern and fossil floras are illustrated and the advantages and disadvantages of the coexistence approach are discussed.

Terrestrial Climate Evolution in Northwest Germany Over the Last 25 Million Years

Abstract The first detailed reconstruction of the continental paleoclimate evolution of the Northwest German Tertiary (Late Oligocene to Pliocene) is presented. The paleoclimate data are derived from the paleobotanical record using the coexistence approach, a method recently introduced that employs climatic requirements of the Nearest Living Relatives of a fossil flora. Twenty six megafloras (fruits and seeds, leaves, woods) from the Tertiary succession of the Lower Rhine Basin and neighboring areas are analyzed with respect to ten meteorological parameters. Additionally, two sample sets from Late Miocene to Early Pliocene sediments comprising 396 palynofloras are analyzed by the same method providing a higher temporal resolution. The temperature curves show a comparatively cooler phase in the Late Oligocene, a warm interval the Middle Miocene, and a cooling starting at 14 Ma. The cooling trend persisted until Late Pliocene with a few higher frequency temperature variations observed. From the beginning of Late Miocene to the present, the seasonality increases and climate appears to have been less stable. As indicated by the precipitation data, a Cfa climate with wet summers persisted in NW Germany from Late Oligocene to Late Pliocene. The results obtained are well in accordance with regional and global isotope curves derived from the marine record, and allow for a refined correlation of the Tertiary succession in the Lower Rhine Basin with the international standard. It is shown that the reconstructed data are largely consistent with the continental climate record for the Northern Hemisphere, as reported by various authors. Discrepancies with previous reconstructions are discussed in detail.

The Miocene peat-forming vegetation of northwestern Germany: an analysis of wood remains and comparison with previous palynological interpretations

New methods of sampling, preparing and identifying fossil wood fragments preserved in browncoal are presented and used to aid in reconstructions of the vegetation from the Miocene Lower Rhine Basin (Germany). The technique involves laboratory charring wood specimens followed by routine identification with reflected light microscopy. Problems concerning the taxonomic identification of the fossil wood are discussed. A list of taxa is presented for the Garzweiler Seam and the implications for reconstructing the Miocene palaeoenvironment are discussed with reference to ecological comparisons of extant taxa: the nearest living relative approach. Results indicate a dominance of conifer wood types, mostly representing members of the Taxodiaceae and Cupressaceae; extensive sampling is required of both small (macroscopic) and large fragments to obtain results comparable with palynological assemblages. Up to now, twenty-one taxa of gymnosperms and angiosperms have been recognized from the Garzweiler seam. Data obtained are compared with those from pollen analysis and corroborate the previously identified trend consisting of increasing concentrations of Taxodium towards the upper section of the seam.

Reconstructing palaeotemperatures using leaf floras – case studies for a comparison of leaf margin analysis and the coexistence approach

Abstract In the past the problems and advantages of the nearest-living-relative (NLR) and leaf physiognomy approaches have been repeatedly discussed and it has been demonstrated that both approaches frequently show broad agreement with each other. However, detailed comparisons of the various methods for accuracy in estimation of palaeoclimate at individual localities are still lacking. Such studies are needed before data obtained from different approaches can be integrated in palaeoclimate maps and models. Moreover, there are some indications that leaf physiognomy and NLR approaches may lead to different results. In this study we applied a physiognomic method based on leaf margin analysis and the coexistence approach, a recent variation of the NLR approach, to two Tertiary palaeofloras (Schrotzburg, Middle Miocene, south Germany; Kleinsaubernitz, Upper Oligocene, east Germany). We demonstrated that both approaches can produce reasonable and consistent results if the standard error of the leaf physiognomy palaeoclimate data is taken into account. However, our results and interpretations indicate that reconstructions based on leaf physiognomy are influenced by factors not related to climate, such as sample size and differential preservation or transport. In contrast, reconstructions for the same fossil assemblages based on the coexistence approach seem to be less affected by taphonomic variables, but may be less sensitive to minor climate changes.

Paleogene evolution of precipitation in northeastern China supporting the middle Eocene intensification of the East Asian monsoon

ABSTRACT The timing of the development of the East Asian monsoon in the geologic past is critically important for paleoclimatological studies, yet few quantitative data are available. Based on palynomorphs from six formations, supplemented by leaf fossils from one of these formations in Fushun, northeastern China, we present a quantitative estimate of the evolution of precipitation in this area during the middle Paleocene–late Eocene. The results demonstrate that seasonal precipitation prevailed during the interval, suggesting that the monsoonal system had already developed by this time. Comparing Paleogene climatic results from different latitudes in eastern China, we conclude that the East Asian monsoon must have been significantly enhanced after the late middle Eocene (∼41–40 Ma), due to increased precipitation differentiation between wet and dry months as shown in the present study. The influence of both the uplift of the Da Hinggan Mountains in northeastern Asia on regional topography and the India-Asia collision globally may have contributed to early monsoon intensification by their influence on air mass movement and associated precipitation patterns in the monsoonal realm.

The Fossil History of Quercus

The evolution of plant ecosystems during the Cenophytic was complex and influenced by both abiotic and biotic factors. Among abiotic forces were tectonics, the distribution of continents and seas, climate, and fires; of biotic factors were herbivores, pests, and intra- and interspecific competition. The genus Quercus L. (Quercoideae, Fagaceae) evolved in this context to become an established member of the plant communities of the Northern Hemisphere, commencing in the Paleogene and spreading to a diverse range of environments in the later Cenozoic. Its palaeontological record, dominated by leaves and pollen, but also including wood, fruits and flowers, is widespread in Eurasia and North America. Consequently, a great number of species have been described, from the 19th century to the present day. Although Quercus is currently an ecologically and economically important component of the forests in many places of the Northern Hemisphere and Southeastern Asia, no comprehensive summary of its fossil record exists. The present work, written by an international team of palaeobotanists, provides the first synthesis of the fossil history of the oaks from their appearance in the early Paleogene to the Quaternary.

Regional-scale palaeoclimate modelling on soft proxy-data basis — an example from the Upper Miocene of the Lower Rhine Embayment

Abstract A new method in palaeoclimatology, initially proposed by Mosbrugger and Schilling (1992) , which makes it possible to extract a maximum of information from the available proxy-data is tested. It possesses a capacity to produce a detailed palaeoclimate reconstruction, which is consistent with both proxy-data and the physics of atmospheric models. Since proxy-data information is particularly poor on a global scale, our method focuses on smaller-scale regions of better proxy-data coverage. It consists of an iterative coupling of inverse mesoscale atmospheric modelling with proxy-based palaeoclimate reconstruction. Using this method, the palaeoclimate of the Tortonian (Upper Miocene) of the Lower Rhine Embayment (northwest Germany) is reconstructed with a high spatial resolution. The results include a physically consistent inter- and extrapolation of proxy-derived palaeoclimate data in space and onto variables not accessible from geological–palaeontological proxies alone. Moreover, the method delivers statistics of the large-scale atmospheric flows as information on the coupling of the regional climate to global fields.

Cenozoic plant diversity of Yunnan: A review

Yunnan in southwestern China is renowned for its high plant diversity. To understand how this modern botanical richness formed, it is critical to investigate the past biodiversity throughout the geological time. In this review, we present a summary on plant diversity, floristics and climates in the Cenozoic of Yunnan and document their changes, by compiling published palaeobotanical sources. Our review demonstrates that thus far a total of 386 fossil species of ferns, gymnosperms and angiosperms belonging to 170 genera within 66 families have been reported from the Cenozoic, particularly the Neogene, of Yunnan. Angiosperms display the highest richness represented by 353 species grouped into 155 genera within 60 families, with Fagaceae, Fabaceae, Lauraceae and Juglandaceae being the most diversified. Most of the families and genera recorded as fossils still occur in Yunnan, but seven genera have disappeared, including Berryophyllum, Cedrelospermum, Cedrus, Palaeocarya, Podocarpium, Sequoia and Wataria. The regional extinction of these genera is commonly referred to an aridification of the dry season associated with Asian monsoon development. Floristic analyses indicate that in the late Miocene, Yunnan had three floristic regions: a northern subtropical floristic region in the northeast, a subtropical floristic region in the east, and a tropical floristic region in the southwest. In the late Pliocene, Yunnan saw two kinds of floristic regions: a subalpine floristic region in the northwest, and two subtropical floristic regions separately in the southwest and the eastern center. These floristic concepts are verified by results from our areal type analyses which suggest that in the Miocene southwestern Yunnan supported the most Pantropic elements, while in the Pliocene southwestern Yunnan had abundant Tropical Asia (Indo–Malaysia) type and East Asia and North America disjunct type that were absent from northwestern Yunnan. From the late Miocene to late Pliocene through to the present, floristic composition and vegetation types changed markedly, presumably in response to altitude changes and coeval global cooling. An integration of palaeoclimate data suggests that during the Neogene Yunnan was warmer and wetter than today. Moreover, northern Yunnan witnessed a pronounced temperature decline, while southern Yunnan experienced only moderate temperature changes. Summer precipitation was consistently higher than winter precipitation, suggesting a rainfall seasonality. This summary on palaeoclimates helps us to understand under what conditions plant diversity occurred and evolved in Yunnan throughout the Cenozoic.

Miocene shift of European atmospheric circulation from trade wind to westerlies

The modern European climatic regime is peculiar, due to its unitary winter but diverse summer climates and a pronounced Mediterranean climate in the south. However, little is known on its evolution in the deep time. Here we reconstruct the European summer climate conditions in the Tortonian (11.62–7.246 Ma) using plant fossil assemblages from 75 well-dated sites across Europe. Our results clearly show that the Tortonian Europe mainly had humid to subhumid summers and no arid climate has been conclusively detected, indicating that the summer-dry Mediterranean-type climate has not yet been established along most of the Mediterranean coast at least by the Tortonian. More importantly, the reconstructed distribution pattern of summer precipitation reveals that the Tortonian European must have largely been controlled by westerlies, resulting in higher precipitation in the west and the lower in the east. The Tortonian westerly wind field appears to differ principally from the trade wind pattern of the preceding Serravallian (13.82–11.62 Ma), recently deduced from herpetofaunal fossils. Such a shift in atmospheric circulation, if ever occurred, might result from the development of ice caps and glaciers in the polar region during the Late Miocene global cooling, the then reorganization of oceanic circulation, and/or the Himalayan-Tibetan uplift.

Facies and paleogeography of the Tertiary of the Lower Rhine Basin - sedimentary versus climatic control

Based on recent studies, the impact of global sea-level and climate change on the paleogeographic and sedimentary evolution of the Tertiary of the Lower Rhine Basin is analysed. It is shown that major changes in global climate and sea-level, such as the high-stand during the Middle Miocene climate optimum, the extreme low-stands near the base of the Tortonian and within the Messinian, are clearly reflected in the sedimentary succession. Continental climate curves, as reconstructed from Tertiary macrofloras of the Lower Rhine Basin, can be correlated with the marine, long-term isotope record. As shown by the analyses, a warm and humid climate with mean annual temperatures above 13 ° C and mean annual precipitation not below 1000 mm, persisted throughout the Late Miocene to Early Pliocene. Continental, high-resolution, climate data show that Late Miocene alternations of lignites and elastics are rather caused by tectonic and sedimentary processes (such as the repeated migration of the river system) than by major climate changes.