Ze’ev Gedalof

Interdecadal climate variability and regime‐scale shifts in Pacific North America

A transect of climate sensitive tree ring-width chronologies from coastal western North America provides a useful proxy index of North Pacic ocean-atmosphere variability since 1600 AD. Here we use this high-resolution record to identify intervals of an enhanced interdecadal cli- mate signal in the North Pacic, and to assess the timing and magnitude of abrupt shifts in this system. In the con- textofthisrecord,thestep-likeclimateshiftthatoccurredin 1976-1977 is not a unique event, with similar events having occurredfrequentlyduringthepast400years. Furthermore, most of the pre-instrumental portion of this record is char- acterized by pronounced interdecadal variability, while the secularportionismorestronglyinterannualinnature. Ifthe 1976-1977 event marks a return to this mode of variability there may be signicant consequences for natural resources management in the North Pacic Sector.

ATMOSPHERIC, CLIMATIC, AND ECOLOGICAL CONTROLS ON EXTREME WILDFIRE YEARS IN THE NORTHWESTERN UNITED STATES

Wildland fire is an important disturbance agent in forests of the American Northwest. Historical fire suppression efforts have contributed to an accumulation of fuels in many Northwestern forests and may result in more frequent and/or more severe wildfire events. Here we investigate the extent to which atmospheric and climatic variability may contribute to variability in annual area burned on 20 National Forests in Washington, Oregon, and Idaho. Empirical orthogonal function (EOF) analysis was used to identify coherent patterns in area burned by wildfire in the Pacific Northwest. Anomaly fields of 500-hPa height were regressed onto the resulting principal-component time series to identify the patterns in atmospheric circulation that are associated with variability in area burned by wildfire. Additionally, cross-correlation functions were calculated for the Palmer drought severity index (PDSI) over the year preceding the wildfire season. Parallel analyses based on superposed epoch analysis focused only on the extreme fire years (both large and small) to discriminate the controls on extreme years from the linear responses identified in the regression analyses. Four distinct patterns in area burned were identified, each associated with distinct climatic processes. Extreme wildfire years are forced at least in part by an- tecedent drought and summertime blocking in the 500-hPa height field. However the re- sponse to these forcings is modulated by the ecology of the dominant forest. In more mesic forest types antecedent drought is a necessary precondition for forests to burn, but it is not a good predictor of area burned due to the rarity of subsequent ignition. At especially dry locations, summertime blocking events can lead to increases in area burned even in the absence of antecedent drought. At particularly xeric locations summertime cyclones can also lead to increased area burned, probably due to dry lightning storms that bring ignition and strong winds but little precipitation. These results suggest that fuels treatments alone may not be effective at reducing area burned under extreme climatic conditions and fur- thermore that anthropogenic climate change may have important implications for forest

Elevation alters ecosystem properties across temperate treelines globally

Temperature is a primary driver of the distribution of biodiversity as well as of ecosystem boundaries. Declining temperature with increasing elevation in montane systems has long been recognized as a major factor shaping plant community biodiversity, metabolic processes, and ecosystem dynamics. Elevational gradients, as thermoclines, also enable prediction of long-term ecological responses to climate warming. One of the most striking manifestations of increasing elevation is the abrupt transitions from forest to treeless alpine tundra. However, whether there are globally consistent above- and belowground responses to these transitions remains an open question. To disentangle the direct and indirect effects of temperature on ecosystem properties, here we evaluate replicate treeline ecotones in seven temperate regions of the world. We find that declining temperatures with increasing elevation did not affect tree leaf nutrient concentrations, but did reduce ground-layer community-weighted plant nitrogen, leading to the strong stoichiometric convergence of ground-layer plant community nitrogen to phosphorus ratios across all regions. Further, elevation-driven changes in plant nutrients were associated with changes in soil organic matter content and quality (carbon to nitrogen ratios) and microbial properties. Combined, our identification of direct and indirect temperature controls over plant communities and soil properties in seven contrasting regions suggests that future warming may disrupt the functional properties of montane ecosystems, particularly where plant community reorganization outpaces treeline advance.

Coherent Region-, Species-, and Frequency-Dependent Local Climate Signals in Northern Hemisphere Tree-Ring Widths*

AbstractPatterns of correlation between tree rings and local temperature or precipitation are investigated using 762 International Tree-Ring Data Bank standardized ring width site chronology time series, and a gridded dataset of temperature and precipitation. Coherent regional- and, in some cases, hemispheric-scale patterns of correlation are found in the extratropical Northern Hemisphere for both the summer prior to and the summer concurrent with ring width formation across different species and over large distances.Among those chronologies that are primarily linked to temperature, thicker ring widths are generally associated with anomalously cool prior summer temperature and anomalously warm concurrent summer temperature. Reconstructions of local summer temperature using prior, concurrent, and/or subsequent year ring widths as predictors demonstrate that useful climate–growth information generally exists in ring widths that are both concurrent with and subsequent to the summer temperature anomaly. Consi...

Climate and Spatial Patterns of Wildfire in North America

Climate interacts with wildfire at a range of spatial and temporal scales. In this chapter I describe a conceptual model that describes how climate (a top-down control) interacts with processes of vegetation development and topography (bottom-up controls) to give rise to characteristic disturbance regimes and observed patterns of wildfire throughout North America. At the shortest timescales (synoptic to seasonal), climate influences fine fuel moisture, ignition frequency, and rates of wildfire spread. At intermediate timescales (annual to interannual), climate affects the relative abundance and continuity of fine fuels, as well as the abundance and moisture content of coarser fuels. At longer timescales (decadal to centennial) climate determines the assemblage of species that can survive at a particular location. Interactions between these species’ characteristics and the influence of climatic processes on wildfire activity give rise to the characteristic disturbance regime and vegetation structure at a given location.

Predicting Potential Global Distributions of Two Miscanthus Grasses: Implications for Horticulture, Biofuel Production, and Biological Invasions

In many regions, large proportions of the naturalized and invasive non-native floras were originally introduced deliberately by humans. Pest risk assessments are now used in many jurisdictions to regulate the importation of species and usually include an estimation of the potential distribution in the import area. Two species of Asian grass (Miscanthus sacchariflorus and M. sinensis) that were originally introduced to North America as ornamental plants have since escaped cultivation. These species and their hybrid offspring are now receiving attention for large-scale production as biofuel crops in North America and elsewhere. We evaluated their potential global climate suitability for cultivation and potential invasion using the niche model CLIMEX and evaluated the models’ sensitivity to the parameter values. We then compared the sensitivity of projections of future climatically suitable area under two climate models and two emissions scenarios. The models indicate that the species have been introduced to most of the potential global climatically suitable areas in the northern but not the southern hemisphere. The more narrowly distributed species (M. sacchariflorus) is more sensitive to changes in model parameters, which could have implications for modelling species of conservation concern. Climate projections indicate likely contractions in potential range in the south, but expansions in the north, particularly in introduced areas where biomass production trials are under way. Climate sensitivity analysis shows that projections differ more between the selected climate change models than between the selected emissions scenarios. Local-scale assessments are required to overlay suitable habitat with climate projections to estimate areas of cultivation potential and invasion risk.

Modelling desertification risk in the north-west of Jordan using geospatial and remote sensing techniques

Remote sensing, climate, and ground data were used within a geographic information system (GIS) to map desertification risk in the north-west of Jordan. The approach was based on modelling wind and water erosion and incorporating the results with a map representing the severity of drought. Water erosion was modelled by the universal soil loss equation, while wind erosion was modelled by a dust emission model. The extent of drought was mapped using the evapotranspiration water stress index (EWSI) which incorporated actual and potential evapotranspiration. Output maps were assessed within GIS in terms of spatial patterns and the degree of correlation with soil surficial properties. Results showed that both topography and soil explained 75% of the variation in water erosion, while soil explained 25% of the variation in wind erosion, which was mainly controlled by natural factors of topography and wind. Analysis of the EWSI map showed that drought risk was dominating most of the rainfed areas. The combined effects of soil erosion and drought were reflected on the desertification risk map. The adoption of these geospatial and remote sensing techniques is, therefore, recommended to map desertification risk in Jordan and in similar arid environments.

Soil properties as constraints to seedling regeneration beyond alpine treelines in the Canadian Rocky Mountains

ABSTRACT Plants growing at the edges of their range limits are expected to be particularly sensitive to changes in precipitation and temperature regimes associated with climatic change. However, non-climatic factors are increasingly recognized as important constraints to species’ range expansions. Therefore, we assessed the effects of soil provenance with respect to the alpine treeline on the germination, growth, and survival of Engelmann spruce (Picea engelmannii) seedlings. Seedlings were grown under controlled conditions in a growth chamber and greenhouse for ninety days in soils collected from four treeline ecotones in the Canadian Rocky Mountains. By controlling seed source and climatic conditions, and eliminating competition and predation, we attribute differences in seedling viability to soil properties that differed across elevation zones and individual treeline sites. Overall, alpine soils originating from beyond the species’ current elevational range were least amenable to growth, and there was some indication of reduced germination and survival in high-elevation soils. Forest soils, which were coarser and more nutrient rich, hosted seedlings with greater above- and below-ground biomass. Thus, the physical and chemical characteristics of alpine soils in our study region may constrain future treeline expansion, underscoring the importance of incorporating soil properties when considering species’ distributions under climate change.

Limited prospects for future alpine treeline advance in the Canadian Rocky Mountains

Treeline advance has occurred throughout the twentieth century in mountainous regions around the world; however, local variation and temporal lags in responses to climate warming indicate that the upper limits of some treelines are not necessarily in climatic equilibrium. These observations suggest that factors other than climate are constraining tree establishment beyond existing treelines. Using a seed addition experiment, we tested the effects of seed availability, predation and microsite limitation on the establishment of two subalpine tree species (Picea engelmannii and Abies lasiocarpa) across four treelines in the Canadian Rocky Mountains. The effect of vegetation removal on seedling growth was also determined, and microclimate conditions were monitored. Establishment limitations observed in the field were placed in context with the effects of soil properties observed in a parallel experiment. The seed addition experiment revealed reduced establishment with increasing elevation, suggesting that although establishment within the treeline ecotone is at least partially seed limited, other constraints are more important beyond the current treeline. The effects of herbivory and microsite availability significantly reduced seedling establishment but were less influential beyond the treeline. Microclimate monitoring revealed that establishment was negatively related to growing season temperatures and positively related to the duration of winter snow cover, counter to the conventional expectation that establishment is limited by low temperatures. Overall, it appears that seedling establishment beyond treeline is predominantly constrained by a combination of high soil surface temperatures during the growing season, reduced winter snowpack and unfavourable soil properties. Our study supports the assertion that seedling establishment in alpine treeline ecotones is simultaneously limited by various climatic and nonclimatic drivers. Together, these factors may limit future treeline advance in the Canadian Rocky Mountains and should be considered when assessing the potential for treeline advance in alpine systems elsewhere.