Christian Ammer

Admixing broadleaved to coniferous tree species: a review on yield, ecological stability and economics

For several reasons the conversion of mono-species into mixed-species forests is presently a major concern of forest management and policy in Central Europe. Although it is possible to show a clear trend in favour of mixed-species forests, private forest owners and some forest economists have often not favoured mixed forests, assuming that they are less profitable. The trend towards mixed forests seems mainly for ecological reasons, while sound economic analysis of mixed forests is still rare. Based on this background the objective of the study is to answer the following four questions: (1) Does the yield of mixed-species forests differ from that of pure forests? (2) Does the mixing of tree species influence the ecological stability of forests? (3) Is the economic value of a mixed forest less than that of a monoculture? (4) How do forest economic models integrate the findings on yield and ecological stability of mixed forests? To answer these questions a literature review was conducted on the possible impacts of mixed-species forests. In comparison to pure stands a greater yield is not necessarily given in mixed stands. Yet, mixed-species stands are better able to compensate disturbances than monocultures. Moreover, there is substantial evidence that mixed-species stands are more resistant against biotic and abiotic disturbances. Applying an extended forest economic model, it was possible to demonstrate that mixing large blocks of native broadleaf species into pure conifer forests may lead to a significant reduction of financial risk. From a risk-averse perspective the economic value of a mixed-species forest may thus be greater than that of a mono-species forest. Yet, it became clear that forest economists do not often integrate the research findings on yield and ecological stability of mixed stands in modelling, but rather apply simple bioeconomic modelling. Moreover, in the context of mixed forests economists also largely ignore even classical financial approaches, which consider risk and risk preferences. We concluded that forest economics has to close substantial research gaps. Firstly, the knowledge of how to integrate biophysical properties of mixed forests in bioeconomic modelling is still an open question. Secondly, forest economists have to adopt the modern approaches of financial theory and management science to value mixed forests.

Impact of ungulates on structure and dynamics of natural regeneration of mixed mountain forests in the Bavarian Alps

Abstract For about the last 20 years, investigations have been carried out to determine the influence of various ecological factors on the natural regeneration of a mixed mountain forest ( Picea abies (L.) Karst, Abies alba Mill., Fagus sylvatica L., Acer pseudoplatanus L.) in the Bavarian Alps. One of these factors is the influence of a ungulate population involving three species ( Capreolus capreolus (L.), Cervus elaphus L., Rupicarpa rupicarpa (L.)). Investigation results show that ungulates play a very important part on the structure and dynamics of the regeneration of this forest. Without browsing, growth rate and species composition of the natural regeneration are mainly determined by the light conditions, which in turn are derived from difference silvicultural treatments. On the shelterwood and the clearcut plots height and height increment also increasingly depend on intra- and interspecific competition effects. The impact of ungulates brings about a complete change in the situation. A high percentage of fir and sycamore saplings is damaged. The survival rates of older fir plants are drastically reduced. In all species, except spruce, height is significantly reduced. A huge loss of estimated biomass was observed. Subsequently the interference processes amongst the saplings also changed. Competition, while still intensive, is no longer important.

Adaptive forest management in central Europe: Climate change impacts, strategies and integrative concept

Abstract Climatic warming may lead to increased or decreased future forest productivity. However, more frequent heat waves, droughts and storms and accompanying pathogen attacks are also expected for Europe and are considered to be increasingly important abiotic and biotic stress factors for forests. Adaptive forestry can help forest ecosystems to adapt to these new conditions in order to achieve management goals, maintain desired forest ecosystem services and reduce the risks of forest degradation. With a focus on central Europe, this paper presents the following management strategies: (1) conservation of forest structures, (2) active adaptation, and (3) passive adaptation. The feasibility and criteria for application of the different strategies are discussed. Forest adaptation may entail the establishment of “neonative” forests, including the use and intermixing of native and non-native tree species as well as non-local tree provenances that may adapt better to future climate conditions. An integrative adaptive management concept is proposed that combines (1) species suitability tests and modelling activities at the international scale, (2) priority mapping of adaptation strategies at the national to regional scale, and (3) implementation at the local scale. To achieve this, an international experimental trial system is required to test suitable adaptive measures throughout Europe and worldwide.

Positive biodiversity-productivity relationship predominant in global forests.

Global biodiversity and productivity The relationship between biodiversity and ecosystem productivity has been explored in detail in herbaceous vegetation, but patterns in forests are far less well understood. Liang et al. have amassed a global forest data set from >770,000 sample plots in 44 countries. A positive and consistent relationship can be discerned between tree diversity and ecosystem productivity at landscape, country, and ecoregion scales. On average, a 10% loss in biodiversity leads to a 3% loss in productivity. This means that the economic value of maintaining biodiversity for the sake of global forest productivity is more than fivefold greater than global conservation costs. Science, this issue p. 196 Global forest inventory records suggest that biodiversity loss would result in a decline in forest productivity worldwide. INTRODUCTION The biodiversity-productivity relationship (BPR; the effect of biodiversity on ecosystem productivity) is foundational to our understanding of the global extinction crisis and its impacts on the functioning of natural ecosystems. The BPR has been a prominent research topic within ecology in recent decades, but it is only recently that we have begun to develop a global perspective. RATIONALE Forests are the most important global repositories of terrestrial biodiversity, but deforestation, forest degradation, climate change, and other factors are threatening approximately one half of tree species worldwide. Although there have been substantial efforts to strengthen the preservation and sustainable use of forest biodiversity throughout the globe, the consequences of this diversity loss pose a major uncertainty for ongoing international forest management and conservation efforts. The forest BPR represents a critical missing link for accurate valuation of global biodiversity and successful integration of biological conservation and socioeconomic development. Until now, there have been limited tree-based diversity experiments, and the forest BPR has only been explored within regional-scale observational studies. Thus, the strength and spatial variability of this relationship remains unexplored at a global scale. RESULTS We explored the effect of tree species richness on tree volume productivity at the global scale using repeated forest inventories from 777,126 permanent sample plots in 44 countries containing more than 30 million trees from 8737 species spanning most of the global terrestrial biomes. Our findings reveal a consistent positive concave-down effect of biodiversity on forest productivity across the world, showing that a continued biodiversity loss would result in an accelerating decline in forest productivity worldwide. The BPR shows considerable geospatial variation across the world. The same percentage of biodiversity loss would lead to a greater relative (that is, percentage) productivity decline in the boreal forests of North America, Northeastern Europe, Central Siberia, East Asia, and scattered regions of South-central Africa and South-central Asia. In the Amazon, West and Southeastern Africa, Southern China, Myanmar, Nepal, and the Malay Archipelago, however, the same percentage of biodiversity loss would lead to greater absolute productivity decline. CONCLUSION Our findings highlight the negative effect of biodiversity loss on forest productivity and the potential benefits from the transition of monocultures to mixed-species stands in forestry practices. The BPR we discover across forest ecosystems worldwide corresponds well with recent theoretical advances, as well as with experimental and observational studies on forest and nonforest ecosystems. On the basis of this relationship, the ongoing species loss in forest ecosystems worldwide could substantially reduce forest productivity and thereby forest carbon absorption rate to compromise the global forest carbon sink. We further estimate that the economic value of biodiversity in maintaining commercial forest productivity alone is

How to quantify forest management intensity in Central European forests

166 billion to

Site adapted admixed tree species reduce drought susceptibility of mature European beech

490 billion per year. Although representing only a small percentage of the total value of biodiversity, this value is two to six times as much as it would cost to effectively implement conservation globally. These results highlight the necessity to reassess biodiversity valuation and the potential benefits of integrating and promoting biological conservation in forest resource management and forestry practices worldwide. Global effect of tree species diversity on forest productivity. Ground-sourced data from 777,126 global forest biodiversity permanent sample plots (dark blue dots, left), which cover a substantial portion of the global forest extent (white), reveal a consistent positive and concave-down biodiversity-productivity relationship across forests worldwide (red line with pink bands representing 95% confidence interval, right). The biodiversity-productivity relationship (BPR) is foundational to our understanding of the global extinction crisis and its impacts on ecosystem functioning. Understanding BPR is critical for the accurate valuation and effective conservation of biodiversity. Using ground-sourced data from 777,126 permanent plots, spanning 44 countries and most terrestrial biomes, we reveal a globally consistent positive concave-down BPR, showing that continued biodiversity loss would result in an accelerating decline in forest productivity worldwide. The value of biodiversity in maintaining commercial forest productivity alone—US

Trends in North American and European regeneration research under the ecosystem management paradigm

166 billion to 490 billion per year according to our estimation—is more than twice what it would cost to implement effective global conservation. This highlights the need for a worldwide reassessment of biodiversity values, forest management strategies, and conservation priorities.

Forest vegetation management under debate: an introduction

Existing approaches for the assessment of forest management intensity lack a widely accepted, purely quantitative measure for ranking a set of forest stands along a gradient of management intensity. We have developed a silvicultural management intensity indicator (SMI) which combines three main characteristics of a given stand: tree species, stand age and aboveground, living and dead wooden biomass. Data on these three factors are used as input to represent the risk of stand loss, which is a function of tree species and stand age, and stand density, which is a function of the silvicultural regime, stand age and tree species. Consequently, the indicator consists of a risk component (SMIr) and a density component (SMId). We used SMI to rank traditional management of the main Central European tree species: Norway spruce (Picea abies [Karst.] L.), European beech (Fagus sylvatica L.), Scots pine (Pinus sylvestris L.), and oak (Quercus robur L. and Quercus petraea L.). By analysing SMI over their whole rotation period, we found the following ranking of management intensity: oak<beech<pine≪spruce. Additionally, we quantified the SMI of actual research plots of the German Biodiversity exploratories, which represent unmanaged and managed forest stands including conifer forests cultivated outside their natural range. SMI not only successfully separate managed from unmanaged forests, but also reflected the variability of forest management and stand properties across the entire sample and within the different management groups. We suggest using SMI to quantify silvicultual management intensity of stands differing in species composition, age, silvicultural system (even-aged vs. uneven-aged), thinning grade and stages of stand conversion from one stand type into another. Using SMI may facilitate the assessment of the impact of forest management intensity on biodiversity in temperate forests.

Biomass functions for the two alien tree species Prunus serotina Ehrh. and Robinia pseudoacacia L. in floodplain forests of Northern Italy

Some forest-related studies on possible effects of climate change conclude that growth potential of European beech (Fagus sylvatica L.) might be impaired by the predicted increase in future serious drought events during the growing season. Other recent research suggests that not only multiyear increment rates but also growth resistance and recovery of beech during, respectively, after dry years may differ between pure and mixed stands. Thus, we combined dendrochronological investigations and wood stable isotope measurements to further investigate the impact of neighborhood diversity on long-term performance, short-term drought response and soil water availability of European beech in three major geographic regions of Germany. During the last four decades, target trees whose competitive neighborhood consisted of co-occurring species exhibited a superior growth performance compared to beeches in pure stands of the same investigation area. This general pattern was also found in exceptional dry years. Although the summer droughts of 1976 and 2003 predominantly caused stronger relative growth declines if target trees were exposed to interspecific competition, with few exceptions they still formed wider annual rings than beeches growing in close-by monocultures. Within the same study region, recovery of standardized beech target tree radial growth was consistently slower in monospecific stands than in the neighborhood of other competitor species. These findings suggest an improved water availability of beech in mixtures what is in line with the results of the stable isotope analysis. Apparently, the magnitude of competitive complementarity determines the growth response of target beech trees in mixtures. Our investigation strongly suggest that the sensitivity of European beech to environmental constrains depends on neighborhood identity. Therefore, the systematic formation of mixed stands tends to be an appropriate silvicultural measure to mitigate the effects of global warming and droughts on growth patterns of Fagus sylvatica.

Response of Fagus sylvatica seedlings to root trenching of overstorey Picea abies

Forest management on many ownerships in North America and Europe has shifted toward the ecosystem management paradigm. The associated shift toward multiple management objectives and focus on natural development patterns should also be reflected in regeneration research efforts. As new information needs arise, research questions and approaches should be evaluated whether they are still appropriate. Specifically, spatial and temporal scales of research studies need to be expanded to accommodate complex sets of management objectives and constraints, rather than being focused on optimal tree regeneration. At the same time, silviculturists are asked to utilize natural trends as a guide for management, but most natural disturbance studies have focused on stand structures and not the regeneration processes. Criteria commonly used to describe disturbance regimes need to be modified to better guide regeneration research efforts under the ecosystem management paradigm.