G. Valleri

Mediterranean fossil whale falls and the adaptation of mollusks to extreme habitats

The hypothesis that sunken carcasses of Mesozoic marine reptiles and Cenozoic whales acted as evolutionary stepping stones to deep-sea reducing habitats is underlain by the question of whether vent-like, chemosymbiotic specialization fi rst evolved at shelf depths. Fossil skeletons of large whales have long been known from ancient shallow-water strata, but they have never been considered as a source of information on ecosystem development. We present a study on a 3 Ma old fossil whale fall and a survey of other Pliocene fossil skeletons to show that the associated biota is dominated by heterotrophs, with subsidiary chemoautotrophs. The taphonomy of the Mediterranean shelf whale falls shows some differences with respect to deep-water studies. Quantitative analyses of abundance data within a large data set on fossil and modern mollusk families confi rm that deep- and shallow-water communities at reducing habitats are composed of a different set of taxa, i.e., specialists occurring only below the shelf break. Mediterranean carcasses sunken in coastal settings do not seem to be favorable for the evolution of whale-fall specialists among the mollusks. The situation reverses as the shelf break is approached.

Sedimentology and high-resolution sequence stratigraphy of the late middle to late Miocene Angostura Formation (western Borbón Basin, northwestern Ecuador)

An integrated analysis of facies and palaeontological content of the middle to late Miocene (c. 12.4–9.1 Ma) Angostura Formation led to the identification of facies associations indicative of shoreface and inner-shelf settings. The Angostura Formation comprises eight high-frequency sequences that are stacked to form a tectonically driven lowstand sequence set. The most complete examples of sequences are bounded by transgressively modified unconformities and exhibit a threefold subdivision: (1) a basal suite of deepening-upward shoreface sediments (transgressive systems tract), including a base-of-cycle shell concentration; (2) a mid-cycle shell bed, consisting of molluscan shells dispersed in a matrix of inner-shelf muddy fine-grained sandstones; (3) a shallowing-upward unit of inner-shelf to lower shoreface sediments almost barren of mollusc fossils (highstand systems tract). Biostratigraphic constraints allowed a reasonable correlation between sequence bounding unconformities and the late middle to late Miocene high-frequency glacio-eustatic changes derived from recent δ18O studies. This correlation has far-reaching implications and leads to the following conclusions: (1) glacio-eustasy in tune with oxygen isotope changes at fourth-order frequency (200 ka–1 Ma duration) may have been the principal factor regulating stratigraphic packaging in the Angostura Formation; (2) these sequences provide an excellent shallow-marine outcrop record of late middle to late Miocene Antarctic glaciations.

FIRST CETACEAN FOSSIL RECORDS FROM ECUADOR, COLLECTED FROM THE MIOCENE OF ESMERALDAS PROVINCE

Cetacean fossils from Ecuador are reported for the first time on the basis of fragmentary remains referred to odontocetes, collected during our investigations of Neogene stratigraphic sequences outcropping along the northern coast (Esmeraldas Province). One specimen was collected near Las Penas village in the Lower-Middle Miocene sediments of the Viche Formation and consists of ear bones and mandibular fragments surely belonging to Ziphiidae. It represents the oldest record of this family in the southeastern Pacific and one of the few records of this family in South America. The other fossil was collected near Rio Verde village in the Upper Miocene sediments of the Angostura Formation and consists of an isolated tooth exhibiting some ziphiid affinities. Pdf

A new vertebrate fossiliferous site from the Late Quaternary at San José on the north coast of Ecuador: preliminary note

A new fossiliferous site is described south of Manta on the north coast of Ecuador. Estuarine sediments overlying Quaternary terraced deposits contain abundant vertebrate remains belonging to the following taxa: Eremotherium cf. laurillardi or E. rusconii, Haplomastodon chimborazi, and Geochelone s.l. On the basis of geological context and the fossil assemblage, a probable Early Holocene age is suggested, although a latest Pleistocene age cannot be ruled out. This discovery will provide crucial new information to enhance knowledge of the geologic and faunistic evolution of the area.

Paleontologic and biostratigraphic observations on the Pliocene of Camarones (Esmeraldas, Ecuador)

Studies on the Onzole Formation outcropping close to the Camarones village (Esmeraldas Province, Northwestern Ecuador) have been carried out. Analyses on foraminifer, otolith and mollusc assemblages have shown an age referable to the Early Pliocene and a bathyal environment. These communities correspond to those of the modern Tropical East Pacific Province. However in the ichthyofauna some taxa, today related to the cool California Current, occur.RiassuntoÈ stato intrapreso uno studio di dettaglio sulle associazioni a foraminiferi, a molluschi e a otoliti della Formazione Onzole, affiorante nei dintorni del villaggio di Camarones (Provincia di Esmeraldas, Ecuador settentrionale). E stato possibile attribuire al Pliocene Inferiore (Zona N 19; Blow, 1969) i sedimenti considerati. Inoltre è stato possibile riconoscere un ambiente di deposizione batiale superiore. Le associazioni riconosciute sono simili a quelle della bioprovincia del Pacifico orientale tropicale, tuttavia la presenza nell’ittiofauna di taxa attualmente legati alla corrente fredda della California, permette di ipotizzare la formazione, durante il Pliocene Inferiore, di una soglia (Panama-Galapagos-Cocos) che avrebbe separato la provincia del Pacifico orientale tropicale dalla provincia californiana.

Relationships between the Sakarya Zone and the Ankara–Erzincan suture (central-northern Turkey): geological and petrographic data from the Ankara–Çankiri, Çorum and Amasya areas

The study was performed in central-northern Anatolia (from Ankara to Amasya) to investigate the relationships of the Sakarya Zone units and the Izmir–Ankara–Erzincan suture (IAES) melange. It reveals that all the Sakarya Zone units are metamorphic and three main tectonostratigraphic units have been distinguished for the first time: the BAA (metasiliciclastic rocks capped by metacarbonates and varicoloured phyllite), the BKC (poly-metamorphic garnet-bearing micaschist and metabasite with a well-preserved relict HP–LT amphibole in a low-amphibolitic to greenschistfacies framework) and the AMC (meta-arkose passing vertically to carbonate–phyllitic alternations and, then, to a thick succession of prevailing acidic to intermediate–basic metavolcanites and volcanicrich metasediments). The BAA and AMC, whose metamorphic frameworks are of Cimmerian age, underlie the Mesozoic carbonate cover sequences (e.g. t2-3, j3–k1) that often show tectonic detachments and slicing. The piling up of the BAA above the HP–LT BKC can be correlated to the tectonic superposition of two similar units (i.e. the Cimmerian Çangaldağ Complex and the Alpine Middle– Upper Cretaceous Domuzdağ Complex, respectively) defined by previous authors in other sectors of the Central Pontides front. The ophiolitic melange generally underlies the Sakarya Zone, but locally (e.g. SE of Amasya) tectonically rests above the latter, probably owing to back-thrusting that occurred during the Tertiary syn-collisional shortenings and the later strike-slip tectonics. We hypothesize that, also in these areas, the Sakarya Zone–IAES consists of a complex tectonic stack of different units, belonging to different palaeogeographic domains and orogenic events (Cimmerian versus Alpine orogenies), but originated within a single long-lived (since Late Triassic to Paleocene/Eocene times), prograding subduction–accretion system in front of the Laurasian continent.

The Scaglia Toscana Formation of the Monti del Chianti: new lithostratigraphic and biostratigraphic data

The Scaglia Toscana Formation (Scisti Policromi Auctt.) is one of the most investigated formation of the Tuscane Nappe. The Formation is widely exposed in the Chianti Mounts and despite the number of studies in this area, some aspects remain poorly known and debated. In this paper new litho- and bio-stratigraphic data from eight key-sections distributed over the entire area are provided and discussed in order to clarify the stratigraphic relationshpis among different lithostratigraphic members, as well as the depositional ages of each member. The Formation deposited in the Cretaceous-Oligocene time interval and it can be subdivided into five lithostratigraphic members: i) the Argilliti di Brolio (wine-red shales with sporadic siliceous calcilutites and rare interbedded cherts); ii) the Marne del Sugame (red and pink marls, calcareous marls and marly limestones with interbedded calcarenitic beds and ruditic lens-shaped bodies including calcareous-siliceous clasts); iii) the Argilliti di Cintoia (grey-green to black shales, locally with manganese-rich siliceous calcilutites and cherts); iv) the Calcareniti di Montegrossi (thin beds of calcilutites and calcarenites with varicoloured shaly-marly interbeds); and v) the Argilliti e Calcareniti di Dudda (alternating thin beds of calcilutites and calcarenites with varicoloured shaly-marly interbeds). These members deposited in a marine environment and have been interpreted as deposited in a turbiditic system, in which shaly and calcareous turbiditic members have been attributed to a basin plain below the CCD, whereas the marls and marly limestones of the Marne del Sugame Member can be settled in a slope/ramp environment above or close to the CCD. Furthermore, the combination of these new data with structural informations coming from literatures allowed to a better paleogeographic reconstruction of the paleobasin. In order to better explain these data, the paper is accompanied by two geological maps realized in the past but never distributed. The two geological maps, at the scale of 1:25,000, cover the whole area from the Cintoia (south of Florence) to the San Gusme (north of Siena) villages.

Late Pliocene fossils of Ecuador and evolution of the Panamic bioprovince after closure of Central American Isthmus

Studies carried on in the last several years allow us to date the Canoa Formation as Late Pliocene. The rich paleontological record (foraminifers, mollusks, and otoliths) allowed us to outline a first articulate picture of the biogeographic relationships in the tropical eastern Pacific during the Plio-Pleistocene. The mollusk fauna shows a Panamic connotation, as the majority (88.7%) of the extant species are present between the Gulf of California and the coasts of Ecuador or northern Peru. Benthic foraminifers assemblages indicate a temperate character with a certain affinity with the Mexican Pacific microfauna. More than 7000 otoliths have been found in the Canoa Formation that are attributed to 105 taxa of 46 fish families. Only 65% of the fossil association is living today along the Ecuadorian coasts, while the association as a whole shows closer relationships with the extant Californian fish community (about 80%). In particular, a conspicuous group of fishes (15 taxa) today spread in the boreal East Pacific (from Oregonian to Californian Province or limited only to the Californian area) has been recorded in the Canoa Formation. We named these taxa “Californian guests.” After the rising of the Central American Isthmus, climatic variations may have caused changes in the pattern of surface currents and (or) activated coastal upwelling cells, supporting the diffusion of a part of the boreal biota toward more southern areas. At the present state of knowledge, both proximal causes seem to be compatible with the data presented.