Hannu Salminen

Spatially dynamic forest management to sustain biodiversity and economic returns

Production of marketed commodities and protection of biodiversity in natural systems often conflict and thus the continuously expanding human needs for more goods and benefits from global ecosystems urgently calls for strategies to resolve this conflict. In this paper, we addressed what is the potential of a forest landscape to simultaneously produce habitats for species and economic returns, and how the conflict between habitat availability and timber production varies among taxa. Secondly, we aimed at revealing an optimal combination of management regimes that maximizes habitat availability for given levels of economic returns. We used multi-objective optimization tools to analyze data from a boreal forest landscape consisting of about 30,000 forest stands simulated 50 years into future. We included seven alternative management regimes, spanning from the recommended intensive forest management regime to complete set-aside of stands (protection), and ten different taxa representing a wide variety of habitat associations and social values. Our results demonstrate it is possible to achieve large improvements in habitat availability with little loss in economic returns. In general, providing dead-wood associated species with more habitats tended to be more expensive than providing requirements for other species. No management regime alone maximized habitat availability for the species, and systematic use of any single management regime resulted in considerable reductions in economic returns. Compared with an optimal combination of management regimes, a consistent application of the recommended management regime would result in 5% reduction in economic returns and up to 270% reduction in habitat availability. Thus, for all taxa a combination of management regimes was required to achieve the optimum. Refraining from silvicultural thinnings on a proportion of stands should be considered as a cost-effective management in commercial forests to reconcile the conflict between economic returns and habitat required by species associated with dead-wood. In general, a viable strategy to maintain biodiversity in production landscapes would be to diversify management regimes. Our results emphasize the importance of careful landscape level forest management planning because optimal combinations of management regimes were taxon-specific. For cost-efficiency, the results call for balanced and correctly targeted strategies among habitat types.

Timing and intensity of precommercial thinning in Pinus sylvestris stands

The effects of timing and intensity of precommercial thinning were studied in three Scots pine artificially regenerated stands on Vaccinium forest sites in southern Finland. A two-level factorial design (3×3) was used in each stand: thinning at dominant height of 3, 6 and 9 m to 1000, 1600 and 2200 stems ha−1. The effects of the treatments were analysed after a period of 23–25 yrs when the dominant height was 14–15 m. Early thinning resulted in the highest standing volume and amount of merchantable wood, and also in slightly accelerated height development. Thinning to 1000 stems ha−1 caused a considerable production loss, but there were no differences between the densities of 1600 and 2200 stems ha−1. Branches became thicker after early thinning, but the differences between the treatments were negligible for crop trees. Crown ratio was lowest as the result of early or moderate thinning (2200 stems ha−1).

The height-increment record of summer temperature extended over the last millennium in Fennoscandia

New data have allowed us to extend a previous height-increment chronology of Scots pine (Pinus sylvestris L.) at the northern Fennoscandian timberline 817 years backwards in time, from 1561 to 745. Our final transfer model accounts for 31% of the dependent instrumental (mean June—August) temperature variance between 1908 and 2007. According to the 1263 yr long summer temperature proxy, the most severe summers were experienced in 1601, 1790 and 782. Correspondingly, the summers of 1689, 885 and 1123 were the most favourable for growth. Two drastic shifts in temperature variability were also found. The twentieth century experienced a multidecadal change as the cold 1905—1914 period was immediately followed by a warm period from 1915 to 1944. An even more prominent shift occurred in the Middle Ages, as the most severe cold spell during 1135—1164 was preceded by the warmest period only a decade earlier, during 1115—1124. The Fourier spectrum of the reconstruction shows significant concentrations of variance around 33.3, 23.3 and 11 years, and between 2.6 and 3.0 years. The wavelet spectrum was able to date several centres of fluctuating periodicities between 745 and 2007. Furthermore, daily temperature records allowed us to define the major growth forcing climatic factor in more detail than in previous response analyses. The mean temperature during a 53 day season from 14 June to 6 August produced the strongest positive growth response (r 2 = 0.36).

A summer temperature proxy from height increment of Scots pine since 1561 at the northern timberline in Fennoscandia

Height increments of 60 Scots pine trees were used to reconstruct mean June—August temperature variability at interannual to decadal scales from 1561 to 2004. Three standardization methods (67%, 33% flexible splines, and a fixed 22 years spline) were compared in building chronologies in order to optimize the frequency response in relation to major climatic forcing factors. The height-growth chronology built using the 33% spline standardization proved to have the most consistent and time-stable relationship with the summer temperatures. Among the monthly precipitation and temperature variables from previous June to current August, previous July shows the highest correlation with height growth. In addition, both previous June and previous August have significant positive correlations. Our final transfer model accounts for 32.5% of the dependent instrumental temperature variance between 1909 and 2004. The Fourier spectra of the height-growth chronology and mean summer temperature are very similar in appearance, both series having peaks at 2.7—3.2 years, 6.7 years and 15.7 years. Thus, the 444 years long summer temperature reconstruction is limited to high and medium frequencies. The coldest three summers in this record were experienced in years 1601, 1790 and 1903. Correspondingly, the summers of 1626, 1689 and 1598 were the warmest. The 1820s experienced the warmest 10-year mean, while the first decade of the twentieth century was the coldest. Among the 14 non-overlapping 30-year periods between 1561 and 1980, the period 1621—1650 was the warmest and the period 1591—1620 the coldest.

Does current summer temperature contribute to the final shoot length on Pinus sylvestris? A case study at the northern conifer timberline

Summary Scots pine (Pinus sylvestris L.) in its natural distribution range is typically classified as a monocyclic species with its predetermined apical growth. It was confirmed by the material sampled in a 50-year-old Scots pine stand from near the northern conifer timberline at Kaamanen, northern Finland that the main controller of the height increment is the mean July temperature of the previous year. Mean monthly temperatures of the current growing season did not correlate significantly with the annual height increment, neither did they improve the fit of the transfer function models predicting annual height increment. This suggests that at least in the northernmost conditions, where the summer is the shortest, the final length of the annual shoot of Scots pine is determined by the summer temperature of the previous year rather than the current year in normal, undisturbed conditions.

Summer temperature affects the ratio of radial and height growth of Scots pine in northern Finland

Abstract• Radial and height growth chronologies from 150-year-old and 50-year old Scots pine stands, both located near to the northern timberline in Laanila, Finland (68° 30′ N, 27° 28′ E), were cross-correlated with each other and with mean temperatures of various temperature periods defined as months, days or growing-degree-days.• The height-growth chronology correlates significantly with radial growth at lags 1 and 2, and radial growth with height growth at lag 2 when the effect of temperature is omitted. On average, low and high growth years represent cool and warm average growing seasons. The summer temperatures (June, July and August) affect most on growth, but height and radial growth do have a different set of effective temperature periods. Furthermore, July temperature variation affects stronger height than radial growth.• Those years with low height/radial-growth ratio, i.e. relatively higher current year’s radial than next year’s height growth, do have lower growth and cooler-than-average July temperature whereas those years with high height/radial-growth ratio do have clearly warmer mid-summer temperatures.Résumé• Les chronologies de croissance en hauteur et de croissance radiale de peuplements de pins sylvestres âgés de 50 et 150 ans, localisés près de la limite septentrionale de la forêt à Laanila en Finlande (68° 30′ N, 27° 28′ E), ont été inter-corrélées entre elles et avec les températures moyennes de différentes périodes de températures définies en terme de mois, de jours ou de degrés jours de période de croissance.• La chronologie de la croissance en hauteur a été significativement corrélée avec la croissance radiale avec un retard de 1 à 2, et la croissance radiale avec la croissance en hauteur avec un retard de 2 quand l’effet de la température n’est pas pris en compte. En moyenne, les années à croissance élevée et faible représentent des saisons de végétation en moyenne chaudes et fraîches. Les températures estivales (juin, juillet, août) affectent fortement la croissance, mais la croissance radiale et la croissance en hauteur ont un jeu différent de périodes de températures efficaces. En outre, la variation de température de juillet influe plus fortement sur la croissance en hauteur que sur la croissance radiale.• Ces années avec un rapport croissance en hauteur/croissance radiale bas, c’est-à-dire une croissance radiale de l’année en cours relativement plus élevée que la croissance en hauteur de l’année suivante, ont une croissance plus faible et une température plus fraîche que la température moyenne de juillet, alors que ces années avec un rapport croissance en hauteur/croissance radiale élevé ont des températures de milieu d’été nettement plus chaudes.

Global warming potentials of stemwood used for energy and materials in Southern Finland: differentiation of impacts based on type of harvest and product lifetime

Wood harvesting in boreal forests typically consists of sequential harvesting operations within a rotation: a few thinnings and a final felling. The aim of this paper is to model differentiated relative global warming potential (GWP) coefficients for stemwood use from different thinnings and final fellings, and correction factors for long‐lived wood products, potentially applicable in life cycle assessment studies. All thinnings and final fellings influence the development of forest carbon stocks. The climate impact of a single harvesting operation is generated in comparison with no harvesting, thus encountering a methodological problem on how to handle the subsequent operations. The dynamic forest stand simulator MOTTI was applied in the modelling of evolution of forest carbon stocks at landscape level in Southern Finland. The landscape‐level approach for climate impact assessment gave results similar to some stand‐level approaches presented in previous literature that included the same forest C pools and also studied the impacts relative to the no‐harvest situation. The climate impacts of stemwood use decreased over time. For energy use, the impacts were higher or similar in the short term and 0–50% lower in the midterm in comparison with an identical amount of fossil CO2. The impacts were to some extent (approximately 20–40%) lower for wood from intermediate thinnings than for wood from final fellings or first thinnings. However, the study reveals that product lifetime has higher relative influence on the climate impacts of wood‐based value chains than whether the stemwood originates from thinnings or final fellings. Although the evolution of future C stocks in unmanaged boreal forests is uncertain, a sensitivity analysis suggests that landscape‐level model results for climate impacts would not be sensitive to the assumptions made on the future evolution of C stocks in unmanaged forest. Energy use of boreal stemwood seems to be far from climate neutral.

Survival and Early Development of Lodgepole Pine

The effect of site fertility, spacing and mode of regeneration on the survival and stand development of lodgepole pine, Scots pine and Norway spruce was studied in a series of experiments comprising 22 study areas in Finland. After 13-14 yrs, lodgepole pine had a mean survival of 68% in planted and 61% in seeded plots, while Norway spruce had the highest (92%) and Scots pine the second highest survival (82%). The survival of planted lodgepole pine was better the wider the spacing. Best survival was achieved on subdry and dry sites, both with planting and with direct seeding. Dominant height was not affected by spacing, but both basal area and volume at the age of 13-14 yrs were significantly higher the denser the spacing. The average difference in the value of the estimated site index H 50 was slightly under 3 m for the superiority of lodgepole compared with Scots pine. Seeding resulted in site indices almost as high as those of lodgepole planting.

Stability of height positions in young naturally regenerated stands of Scots pine

Abstract The stability of height positions of saplings in young stands of Scots pine (Pinus sylvestris L.) was examined by comparing the height order of trees at successive measurements. The data consisted of six naturally regenerated stands and one planted stand for comparison. The length of the examination period was approximately 20 years, covering the mean height development of stands from 0.5 to 5.5 m. For each stand, a structural equation model was fitted by which the covariance structure occurring in the data could be described. In terms of the stability coefficients between the variables produced by the model, the height position of the saplings was very stable. The height positions had mainly been established during the first 5–10 years of age and at a mean height of less than 0.5–1 m. In the naturally regenerated stands, stability was virtually complete from the stage of 15–20 years and 1.5–2 m onwards. In the planted stand, stability became complete slightly later than in the naturally regenerated stands. According to the results, the height of the sapling can be emphasized as a selection criterion in precommercial thinning.

Dendroclimatological analysis of seeded and thinned Scots pine (Pinus sylvestris L.) stands at the coniferous timberline

Radial growth was examined in two Scots pine stands that were seeded during the 1920s and 1930s due to reforestation and afforestation activity on the timberline of northern Finnish Lapland. Tree-rings of seeded pines were calibrated against the instrumental records of local weather and large-scale atmospheric patterns and further compared to pines of natural origin on the timberline. The studied stands were shown to contain common growth variability that differed from the variability of natural pines. Deviating growth of seeded pines was attributed to their only moderate growth dependence on mid-summer (July) temperatures and, likewise, their strong dependence on the autumn climate in the previous year, and linked to the different genetic origin of artificial stands due to southern seeds. Stands were thinned for different densities in 1985 and 1986. The growth response to thinning was markedly better than could be expected according to previous studies. We found that the positive growth reaction conceivably benefited from the ameliorated winter conditions, expressed as warmed (March) temperatures and the prolonged positive phase of the Arctic Oscillation (February). The results emphasize the determining influence of climatic fluctuations on reforestation and afforestation near the distributional limits of tree species.