Josep Vallverdú

A new early Pleistocene hominin mandible from Atapuerca-TD6, Spain

We present the description of a new mandibular specimen, ATD6-113, recovered in 2006 from the TD6 level of the Gran Dolina cave site in Sierra de Atapuerca, northern Spain. A detailed study of the lithostratigraphy of the top sequence of this level, the section from where all human remains have been recovered so far, is also presented. We have observed that the hominin stratum, previously defined as Aurora Stratum, represents a condensed deposit of at least six layers, which could not be distinguished in the test pit made in 1994-95. Therefore, the human fossil remains were probably deposited during a discrete and undetermined time period. The new mandibular fragment exhibits a very similar morphology to that of the most complete specimen, ATD6-96, which was recovered in 2003 from a different layer. This suggests that both specimens represent the same biological population. The two mandibles, as well as the small mandibular fragment ATD6-5 (which constitutes part of the holotype of Homo antecessor), present a morphological pattern clearly derived with regard to that of the African early Homo specimens usually included in H. habilis and H. rudolfensis, the mandibles D211 and D2735 from Dmanisi, and most of the early Pleistocene mandibles from Sangiran. The TD6 mandibles also exhibit some derived features with regard to the African early Pleistocene specimens included in H. ergaster (or African H. erectus). Thus, the TD6 hominins seem to represent a lineage different from other African and Asian lineages, although some (metric in particular) similarities with Chinese middle Pleistocene mandibles are noted. Interestingly, none of the apomorphic mandibular features of the European middle and early late Pleistocene hominins are present in the TD6 mandibles.

Age and Date for Early Arrival of the Acheulian in Europe (Barranc de la Boella, la Canonja, Spain)

The first arrivals of hominin populations into Eurasia during the Early Pleistocene are currently considered to have occurred as short and poorly dated biological dispersions. Questions as to the tempo and mode of these early prehistoric settlements have given rise to debates concerning the taxonomic significance of the lithic assemblages, as trace fossils, and the geographical distribution of the technological traditions found in the Lower Palaeolithic record. Here, we report on the Barranc de la Boella site which has yielded a lithic assemblage dating to ∼1 million years ago that includes large cutting tools (LCT). We argue that distinct technological traditions coexisted in the Iberian archaeological repertoires of the late Early Pleistocene age in a similar way to the earliest sub-Saharan African artefact assemblages. These differences between stone tool assemblages may be attributed to the different chronologies of hominin dispersal events. The archaeological record of Barranc de la Boella completes the geographical distribution of LCT assemblages across southern Eurasia during the EMPT (Early-Middle Pleistocene Transition, circa 942 to 641 kyr). Up to now, chronology of the earliest European LCT assemblages is based on the abundant Palaeolithic record found in terrace river sequences which have been dated to the end of the EMPT and later. However, the findings at Barranc de la Boella suggest that early LCT lithic assemblages appeared in the SW of Europe during earlier hominin dispersal episodes before the definitive colonization of temperate Eurasia took place.

The lithic industry of Sima del Elefante (Atapuerca, Burgos, Spain) in the context of Early and Middle Pleistocene technology in Europe.

This paper presents the lithic assemblages documented at Sima del Elefante (TE) and their importance in the context of the Early and Middle Pleistocene human occupation of Europe. We also study changes in human behaviour within the context of the palaeoenvironmental evolution of the Sierra de Atapuerca. This site has characteristics that are of great value for the study of human evolution. The lower levels of TE (Units TE7-TE14) are an essential reference for understanding the early stages of the colonization of Europe. The TE9c level has provided stone tools (Mode 1), faunal remains, and human fossils dated to 1.22xa0Ma (millions of years ago). Moreover, this is one of the few European sites with a stratigraphic sequence that includes remains of human occupations predating the Jaramillo subchron (Early Pleistocene) and from the Late Middle Pleistocene (Units TE18-TE19). Despite this, the presence of archaeologically sterile units (TE15-17) prevents us from establishing a continuous relationship between the Early and Middle Pleistocene human settlements and, consequently, between their technological and behavioural differences. We can, however compare the technological and palaeoeconomic strategies adopted by different species of hominins during two key phases of the occupation of Europe.

Les sédiments d'Homo Antecessor de Gran Dolina, (Sierra de Atapuerca, Burgos, Espagne). Interprétation micromorphologique des processus de formation et enregistrement paléoenvironnemental des sédiments

Resume Letude des sediments, a laide de lames minces de grand format, est la continuation dune approche contextuelle integree aux restes dHomo antecessor de TD6 a Gran Dolina. Lapproche contextuelle a debute avec letude archeostratigraphique des restes humains. Letude archeostratigraphique confrontee a lanalyse sedimentaire explique les difficultes de separer les restes humains des autres occupations anthropiques dune partie de TD6, Aurora Stratum archeostratigraphique (ASa). En ce sens, ASa se trouve dans un contexte sedimentaire microstratifie, indiquant une possible synchronie sedimentaire des restes humains, de la couche Aurora Stratum (AS). Lanalyse microscopique des sediments non perturbes dASa montre que la sedimentation est hydrique, a basse energie et denfouissement rapide. Linterpretation paleoenvironnementale est basee sur lenregistrement des processus pedosedimentaires de Gran Dolina. Les rapides changements environnementaux enregistres dans les sediments aident a proposer une interpretation chronoclimatique. Les occupations anthropiques dASa et AS de TD6 a Gran Dolina ont eu lieu pendant une periode glaciaire du Pleistocene inferieur final.

Is it possible to identify temporal differences among combustion features in Middle Palaeolithic palimpsests? The archaeomagnetic evidence: A case study from level O at the Abric Romaní rock-shelter (Capellades, Spain)

Abstract Archaeomagnetic dating is probably one of the most known applications of magnetic methods to archaeology but there are others still underutilized and of particular interest to Palaeolithic archaeology. Here, we report a novel application of archaeomagnetism as a technique to determine temporal diachronies among combustion features from the same surface within palaeolithic palimpsests. The approach is based on the subtle directional changes of the Earths magnetic field through time (secular variation, SV) and on the ability of burned materials to record such variations under certain conditions. Three Middle Palaeolithic hearths from level O (ca. 55xa0kaxa0BP) at the Abric Romani rock-shelter (NE Spain), were archaeomagnetically investigated. The studied surface (black homogeneous carbonaceous facies), recorded the magnetic enhancement produced by fire with a tenfold increase in concentration-dependent magnetic parameters in the uppermost centimetre with respect to its unburned or deeper counterparts. Pseudo-single domain (PSD) Ti-low titanomagnetite was identified as the main remanence carrier. The irreversibility of thermomagnetic curves suggests that these samples did not undergo enough high temperatures as to record a full thermoremanence (TRM). Additionally, the occasional occurrence of maghaemitized magnetite is interpreted as an indication of a thermochemical remanent magnetization (TCRM), making these samples unsuitable for absolute palaeointensity determinations. Two well-defined (α95

The TD6.3 faunal assemblage of the Gran Dolina site (Atapuerca, Spain): a late Early Pleistocene hyena den

Abstract Ethological studies have shown that besides human groups, large-medium carnivores have bone-collecting habits. The research developed since the last half of the twentieth century has attempted to characterise the carnivore’s accumulations and to identify them in the archaeo-paleontological record. At present, we have diagnostic criteria that define the accumulations produced by hyenids (mainly, Crocuta crocuta), thereby allowing us to differentiate them from the other accumulating agents. The faunal assemblage recovered at the Early Pleistocene TD6.3 layer of the Gran Dolina site is characterised by the presence of typical elements described in hyena dens: presence of small carnivores remains, high bone breakage, low epiphysis survival and a high frequency (>30%) of specimens with carnivore induced modifications, including large amounts of digested bones. However, attritional mortality profiles, hyena’s cubs remains, mid-shaft bone cylinders or differential anatomical composition among different weight-sizes, have not been observed or are ambiguous. In addition, anthropic evidences are scarce and concentrated in the uppermost section of the layer. TD6.3 is the result of an accumulation produced by hyenas using the cave as a den, in alternation with sporadic occupations by hominin groups. TD6.3 shows that hyena fossil accumulations may present great anatomical and taxonomic variability.

A Reply to Otterbein

Institut Catala de Paleoecologia Humana i Evolucio Social (IPHES), C/Escorxador s/n, 43003 Tarragona, Spain; Area de Prehistoria, Universitat Rovira i Virgili (URV), Avinguda de Catalunya 35, 43002 Tarragona, Spain (Carbonell, Caceres, Lozano, Saladie, Rosell, Lorenzo, Vallverdu, Huguet, Canals) (icaceresprehistoria.urv.cat)/Visiting Professor, Institute of Vertebrate Paleontology and Paleoanthropology of Beijing (IVPP; Carbonell)/Centro Nacional de Investigacion sobre Evolucion Humana (CENIEH), Paseo Sierra de Atapuerca, s/n 09002, Burgos, Spain (Bermudez de Castro). 15 II 11