Paula J. Noble

Late Ordovician mass extinction: A new perspective from stratigraphic sections in central Nevada

Integrated sequence stratigraphic, biostratigraphic, and chemostratigraphic analyses of three stratigraphic sections in central Nevada indicate that Late Ordovician glaciation-induced sea-level fall produced diachronous, stepwise faunal turnover in graptolites, conodonts, chitinozoans, and radiolarians, and also triggered a strong, but transient, positive δ13C excursion. This pattern is very different from that described for most mass extinction events.

Early Paleozoic radiolarian biozonation

Current knowledge of early Paleozoic radiolarians permits the recognition of faunal assemblages and zones that constitute a biozonation. This new zonation utilizes robust and relatively common, easily recognized morphotypes. It will facilitate rapid acquisition of previously unavailable age constraints for marine siliceous sedimentary successions in hitherto poorly understood early Paleozoic orogenic collages.

The lundgreni Extinction Event: Integration of paleontological and geochemical data from Arctic Canada

Abstract The lundgreni Extinction Event (LEE) in Arctic Canada, as in other regions globally, exhibited an apparently abrupt and catastrophic reduction of graptolite and radiolarian taxa, followed by a short period of extremely low diversity, the dubius-nassa Biozone, followed in the subsequent praedeubeli-deubeli Biochron by the rapid evolutionary diversification of new taxa. The extinction event was very close to, or coincided with, a regression, as well as with a moderately strong positive δ13Corganic excursion. The excursion is bifurcated with the lower fork occurring at or near the LEE and the upper fork occurring in about the lower part of the praedeubeli-deubeli Biozone, possibly coinciding with a modest transgression. Radiolarians are diverse and abundant throughout the lundgreni Biozone, sparse to absent directly following the LEE (i.e., in the dubius-nassa Biozone), and found only sporadically within the praedeubeli-deubeli Biozone. Acritarch frequency throughout the entire investigated interval is extremely low, and only in one level in the lundgreni Biozone so-called “giant acritarchs” were observed. Sphaeromorph acritarchs and amorphous organic matter are moderately common in the lundgreni Biozone, extremely abundant in the brief time interval of the dubius-nassa Biozone at Rookery Creek, and only moderately common in higher strata. The presence of dark yellow and light brown-colored organic matter indicates that the absence of acritarchs is not due to reworking or thermal degradation but is, instead, suggestive of paleoecological control. Proximity to a shallower water shelf may have strongly influenced palynomorph/organic content.

Isotope Geochemistry and Plankton Response to the Ireviken (Earliest Wenlock) and Cyrtograptus lundgreni Extinction Events, Cape Phillips Formation, Arctic Canada

Several Canadian Arctic Silurian stratigraphic sections from the basinal facies of Cape Phillips Formation have been sampled across the Llandovery–Wenlock and early–late Homerian (late Wenlock) boundary intervals for integration of biotal (graptolite, radiolarian, palynomorph) and geochemical (13C) data for two well-known extinction events, the Ireviken and lundgreni Extinction (LEE) events. Graptolites, abundant and well preserved, provide a refined biostratigraphic base for other paleontologic and geochemical data. They were globally affected by both extinction events: about 64% reduction for the Ireviken and 90–95% for the LEE. Recovery from the LEE event was slow and diversity low through the late Homerian. Radiolarians—diverse (28 species), abundant, and beautifully preserved through the early Homerian―are sharply reduced slightly below the LEE boundary. Data for the late Homerian are more scattered, but it appears that diversity was low; few early Homerian taxa crossed the extinction boundary and new taxa appeared. Palynological studies around the LEE interval are at a preliminary level, but it appears that chitinozoans and microflora (acanthomorph acritarchs, prasinophytes, sphaeromorphs) were impacted by the extinction event. Chitinozoans, though seldom abundant, appear to disappear briefly across the LEE boundary, as do palynomorphs. Amorphous organic matter is abundant in the upper part of the lundgreni Zone; it is much less common in the early and middle–late Homerian and common in the latest part. Stable isotope geochemistry shows well-marked, positive excursions in the δ13Corganic fraction associated with the Ireviken event and LEE. The Ireviken excursion (C1) curve has a sharp base, reaches a peak in the early Wenlock, and then tapers more slowly. The LEE excursion (C4) peaks at, or slightly below, the early–late Homerian boundary. Both are positive excursions. Considering the limits of biostratigraphic placement of the boundaries, they were close to or coincident with regressions, particularly across the LEE interval. The δ13Corganic excursions are greater for inshore sections compared with the offshore section. The most parsimonious explanation for increased carbon content is accelerated weathering of carbonates exposed during a lowstand.

Recognition of fine-scale imbricate thrusts in lower Paleozoic orogenic belts—An example from the Roberts Mountains allochthon, Nevada

Dating of lower Paleozoic cherts is now possible through advancements in radiolarian biostratigraphy, permitting greater refinement of structural and stratigraphic models than has previously been possible. A detailed study of the type section of the Ordovician Vinini Formation, Vinini Creek, Roberts Mountains allochthon, Nevada, has yielded anomalously young Devonian ages. These ages were unanticipated and indicate that structural imbrication exists in a classic and previously well dated section of basinal strata. Previous ages of cherts were inferred from their stratigraphic position relative to graptolite-bearing shales. New mapping and biostratigraphy reveal considerable structural interleaving of Ordovician and Devonian strata within the allochthon, and many of the cherts appear to have structural bases. The Roberts Mountains allochthon, considered to be allochthonous but not far traveled, displays a highly complex structural history, including out of sequence thrusting and/or overprinting by later contractional deformation having a similar transport direction. This deformation was previously unrecognized because of the lack of overlap sequences in critical areas and the paucity of internal age control. Because the Roberts Mountains allochthon is not considered to be exotic, but rather has undergone only a modest amount of transport, this study shows that convergent margins with high degrees of structural imbrication do not require large amounts of transport. A biostratigraphic approach that examines all lithologies, coupled with detailed observations of field relations, is required for the construction of detailed and accurate structure sections in early Paleozoic orogenic belts.

KATIAN (ORDOVICIAN) RADIOLARIANS FROM THE MALONGULLI FORMATION, NEW SOUTH WALES, AUSTRALIA, A REEXAMINATION

Abstract Upper Ordovician radiolarians are described from limestone within the Malongulli Formation in the Cliefden Caves area using revised systematics. Eight genera and 12 species are described, including the following four new species: Protoceratoikiscum crossingi, Haplotaeniatum ovatum, Haplotaeniatum prolatum, and Borisella dunhilli, and two new combinations, Borisella subulata and Haplotaeniatum spongium. All samples are dominated by Kalimnasphaera maculosa, B. subulata, and Palaeoephippium octaramosum with lesser abundances of Haplotaeniaum spp. and Inanigutta complanata, and rare occurrences of Protoceratoikicsum spp. The rare species Etymalbaillella yennienii is reported for the first time outside of China. The Malongulli Formation is upper Eastonian through lower Bolindian (Katian global stage) based on graptolite and conodont biostratigraphy. From a stratigraphic standpoint, the limestone intervals sampled appear to represent two or possibly three stratigraphically separate intervals within the Malongulli Formation. Conodonts recovered from Coppermine Creek, Angullong/Belubula, and Gleesons Creek localities occur in the lower and middle parts of the formation and fall within one conodont zone, the O. velicuspis Zone. The stratigraphic position of the limestone at the Sugarloaf Creek locality is higher, possibly as high as lower Bolindian, based on proximity to the top of the formation where Bolindian graptolites have been recovered. Despite the different stratigraphic positions of the limestones, the radiolarians contained within are essentially the same fauna, barring some differences in relative abundance in Haplotaeniatum and Palaeoephippium. The Malongulli fauna is discussed with respect to other Upper Ordovician faunas, most notably those from siliceous facies in the eastern Lachlan Orogen and from the Hanson Creek Formation of Nevada, U.S. The radiolarian species occur across different depositional environments and from a range of geographically widespread localities, which bodes well for them being recognized as biostratigraphically important faunal markers with correlation potential within the Katian.

Middle to Upper Tournasian Radiolaria of the Baltalimani Formation, Istanbul, Turkey

Abstract Radiolarians collected from five localities of the Baltalimani Formation in the Istanbul region are described in detail, and the local stratigraphic ranges and persistence of biostratigraphically important radiolarian species are documented. The spumellarian component is published for the first time, and four new species are described: Albaillella kayai n. sp., Ceratoikiscum jucundum n. sp., Meschedea akcetensis n. sp., and Stigmosphaerostylus istanbulensis n. sp. Two assemblages are recognized: the A. paradoxa and the A. sp. aff. A. undulata-A. deflandrei assemblage. The older A. paradoxa assemblage contains Traenosphaera sicarius, Pararchocyrtium serrensis, Cyrtisphaeractenium mendax, and common to abundant A. paradoxa and Popofskyellium. Albaillella paradoxa is not restricted to the older assemblage but is the only species of Albaillella present. The younger A. sp. aff. A. undulata-A. deflandrei assemblage contains either A. sp. aff. A. undulata or A. deflandrei. Albaillella indensis ambigua, A. kayai n. sp., and Kantollium sp. A occur exclusively in the A. sp. aff. A. undulata-A. deflandrei assemblage, but are rare and patchy in their occurrence. Distinctive taxa common to both assemblages are Archocyrtium lagabriellei, Pylentonema antiqua, C. avimexpectens, C. umbraculatum, C. jucundum n. sp., Trilonche palimbola, M. akcetensis n. sp., S. tortispina, Polyentactinia polygonia, P. aranea, and Cubaxonium? octaedrospongiosum. Many of the taxa present occur in middle and upper Tournasian sections from France, the central Pyrenees, and Germany and provide a solid basis for re-assessing the age of the Baltalimani Formation as middle to late Tournasian (Tn2–Tn3).

Paleoseismic history of the Fallen Leaf segment of the West Tahoe–Dollar Point fault reconstructed from slide deposits in the Lake Tahoe Basin, California-Nevada

The West Tahoe–Dollar Point fault (WTDPF) extends along the western margin of the Lake Tahoe Basin (northern Sierra Nevada, western United States) and is characterized as its most hazardous fault. Fallen Leaf Lake, Cascade Lake, and Emerald Bay are three subbasins of the Lake Tahoe Basin, located south of Lake Tahoe, and provide an opportunity to image primary earthquake deformation along the WTDPF and associated landslide deposits. Here we present results from high-resolution seismic Chirp (compressed high intensity radar pulse) surveys in Fallen Leaf Lake and Cascade Lake, multibeam bathymetry coverage of Fallen Leaf Lake, onshore Lidar (light detection and ranging) data for the southern Lake Tahoe Basin, and radiocarbon dates from piston cores in Fallen Leaf Lake and Emerald Bay. Slide deposits imaged beneath Fallen Leaf Lake appear to be synchronous with slides in Lake Tahoe, Emerald Bay, and Cascade Lake. The temporal correlation of slides between multiple basins suggests triggering by earthquakes on the WTDPF system. If this correlation is correct, we postulate a recurrence interval of ∼3–4 k.y. for large earthquakes on the Fallen Leaf Lake segment of the WTDPF, and the time since the most recent event (∼4.5 k.y. ago) exceeds this recurrence time. In addition, Chirp data beneath Cascade Lake image strands of the WTDPF offsetting the lake floor as much as ∼7.5 m. The Cascade Lake data combined with onshore Lidar allow us to map the WTDPF continuously between Fallen Leaf Lake and Cascade Lake. This improved mapping of the WTDPF reveals the fault geometry and architecture south of Lake Tahoe and improves the geohazard assessment of the region.

UPPER WENLOCK CERATOIKISCIDAE (RADIOLARIA) FROM THE CAPE PHILLIPS FORMATION, ARCTIC CANADA

Abstract Taxa belonging to the Ceratoikiscidae are described in detail from two measured sections of Member C of the Cape Phillips Formation exposed along Rookery Creek, Cornwallis Island, Arctic Canada. The sections encompass uppermost Sheinwoodian (top lower Wenlock) to the base of the Gorstian (base Ludlow) and are dated by graptolites assigned to the Cyrtograptus perneri–Monograptus opimus Zone (upper Sheinwoodian), the Cyrtograptus lundgreni Zone (lower Homerian), and the Colonograptus praedeubeli–Colonograptus deubeli Zone (upper Homerian). Radiolarians are found in great abundance throughout the lower Homerian, allowing for the establishment of the first appearance of the genus Helenifore Ormiston and Nazarov. Helenifore is emended to accommodate the Silurian species H. quadrispina n. sp. and H. speciosus (Furutani 1990). One new genus, Kappaforma, including the new species, K. insecta, is described, as is the new species H. quadrispina. Ceratoikiscum octapleura Renz and C. armiger Furutani are synonymized with C. lenoides Renz, and H. planus Umeda is synonymized with H. speciosus.

Biostratigraphical and palaeobiogeographical implications of Lower Silurian Radiolaria from black cherts of the Armorican Massif (France)

A moderately well-preserved radiolarian assemblage was recovered from organic-rich black chert in a Llandovery (Lower Silurian) sequence that crops out in southern Brittany (Chalonnes-sur-Loire section, Armorican Massif, France). The assemblage is composed of two families (Rotasphaeridae and Haplotaeniatidae), four genera (?Diparvapila, Secuicollacta, Orbiculopylorum, Haplotaeniatum) and 13 species. Some were identified from whole specimens preserved in silica and extracted following dilute hydrofluoric acid processing, while others were recognized in thin-section preparations, as they are beautifully preserved as ‘carbonized’ microfossils. The age range suggested by conodonts and chitinozoans yielded after HF processing from one of the 27 studied samples is in good agreement with the previously published age based on graptolites. The recovered radiolarians are discussed, documented and compared with known Rhuddanian, Aeronian and lower Telychian assemblages in the literature. The stratigraphic ranges are extended for the species Secuicollacta bipola, S. hexactinia, S. parvitesta, Orbiculopylorum granti and O. splendens based on our new data. A significant number of radiolarians found in our samples occur in Llandovery sections from Alaska, Nevada, Arctic Canada and Sweden. These similarities are used to discuss the palaeodistribution of Lower Silurian Radiolaria and our observations support the hypothesis of a wide geographical distribution for these Palaeozoic species.