Ricardo Russo Siewert

The ``Taygetis ypthima species group{''} (Lepidoptera, Nymphalidae, Satyrinae): taxonomy, variation and description of a new species

Abstract A new species of Taygetis Hübner, [1819] (Lepidoptera, Nymphalidae, Satyrinae) from southeastern Brazil is described: Taygetis drogoni sp. n. In addition, T. servius Weymer, 1910 and T. fulginia d’Almeida, 1922 are resurrected from synonymy and a taxonomic discussion on the species T. ypthima Hübner, [1821] and T. rectifascia Weymer, 1907 is provided. A dichotomous key for the species is also provided.

A new species of Magneuptychia Forster, 1964 (Lepidoptera: Nymphalidae: Satyrinae) from Brazilian Savanna

A new species of Satyrinae butterfly from the Brazilian Savanna, Magneuptychia flavofascia n. sp., is described based on adult morphological characters with a discussion about its placement within the genus.

Revision of Godartiana Forster (Lepidoptera: Nymphalidae), with the description of a new species from northeastern Brazil

A revision of Godartiana Forster, 1964 (Satyrinae: Euptychiina) is provided, including the description of a new species, G. amadoi Paluch, Zacca & Freitas sp. nov. from Bahia and Pernambuco, northeastern Brazil. Two genera, Cercyeuptychia L. Miller & Emmel, 1971 syn. nov. and Praefaunula Forster, 1964 syn. nov., are herein synonymised with Godartiana, based on morphological and molecular data from mitochondrial gene Cytochrome C Oxidase I and the nuclear genes GAPDH and RpS5. A new taxonomic arrangement is proposed for Godartiana, increasing to five the number of species known in this genus and also synonymising Euptychia liturata Butler, 1867 syn. nov. and E. vesper Butler, 1867 syn. nov. with G. armilla (Butler, 1867) comb. nov. The female genitalia of all species of Godartiana are described and illustrated, information is given on host plants and biology, and the previously unknown immature stages of G. muscosa are described and illustrated.

A new species of Aricoris Westwood, [1851] belonging to “ chilensis ” group (Lepidoptera: Riodinidae)

A new species of Riodinidae from the grasslands of South Brazil, Paraguay and Argentina, Aricoris schneideri sp. nov., is described.

A new subspecies of Pierella lena (Lepidoptera: Nymphalidae: Satyrinae) in northeastern Brazil

Abstract Pierella lena atlantica Paluch, Zacca & Siewert ssp. nov. (Lepidoptera: Nymphalidae: Satyrinae) is described from the Atlantic Forest of Bahia (Itanagra, Una, and Cachoeira), Brazil. A discussion about this new taxon and its relationship with the other subspecies of P. lena (Linnaeus, 1767) are provided, as are the first illustrations of the female genitalia and an identification key to adults of Pierella hyalinus (Gmelin, [1790]) and P. lena from South America.

Hidden in plain sight: a morphological study revealing three new species of the skipper genus Drephalys Watson, 1893 (Lepidoptera: Hesperiidae) from Brazil

Three new species of Drephalys Watson, 1893 from Brazil are described as follows: D. dracarys Madruga, Siewert, Mielke Dolibaina, sp. n. from Acre, Mato Grosso and Rondônia states, D. electrinus Siewert, Madruga, Mielke Dolibaina, sp. n. from Acre and Amazonas states, and D. citrinus Madruga, Siewert, Mielke Casagrande, sp. n. from Amazonas state. Drephalys heraclides Bell, 1942 was reported for the first time from Brazil. To characterize the new species, illustrations of the male genitalia of D. heraclides, D. phoenice (Hewitson, 1867) and D. phoenicoides (Mabille Boullet, 1919) are provided. Diagnosis, adult photos, illustration of the male genitalia and a geographical distribution map are provided for the new species.

Redescription of Yphthimoides patricia (Hayward, 1957), with taxonomic notes on the names Euptychia saltuensis Hayward, 1962 and Yphthimoides manasses (C. Felder & R. Felder, 1867) (Nymphalidae: Satyrinae)

Euptychia saltuensis Hayward, 1962, new synonym, currently regarded as a nomen dubium and possibly a junior subjective synonym of Yphthimoides manasses (C. Felder R. Felder, 1867), is here treated as a junior subjective synonym of Yphthimoides patricia (Hayward, 1957), based on morphological characters of the male genitalia and the DNA barcode. The taxonomic status of Y. patricia is re-examined, and a detailed redescription of the adult morphology, including the male genitalia, is presented. Information on the distribution, habitat and immature stages of Y. patricia is also provided. Yphthimoides patricia is clearly a distinct species from Y. manasses based on the analysis of DNA barcode sequences and the morphology of the male genitalia.

Mesene Doubleday (Lepidoptera: Riodinidae) from Parque Nacional da Serra do Divisor, Acre, Brazil, with taxonomic notes and the description of a new species

The species of Mesene Doubleday, 1847 collected at the Parque Nacional da Serra do Divisor, Acre, Brazil (PNSD) are listed and supplemented by taxonomic and behavioral notes. In total, 30 specimens were collected, representing seven species including one new: M. monostigma monostigma (Erichson, [1849]), M. nepticula stigmosa Stichel, 1910, M. eupteryx Bates, 1868 stat. rest., M. paraena Bates, 1868, M. epaphus pyrrha Bates, 1868, M. leucophrys Bates, 1868, and M. gracielae sp. nov. The following notes on the taxonomy of Mesene are proposed: M. eupteryx Bates, 1868 stat. rest. is recognized as a valid species, M. epaphus sertata Stichel, 1910 syn. nov. is a new synonym of M. epaphus pyrrha Bates, 1868, and M. lecointrei P. Jauffret & J. Jauffret, 2008 syn. nov. is a new synonym of M. philonis Hewitson, 1874. Mesene margaretta (White, 1843), M. philonis and M. nola are recognized as closely related. Species encountered at the PNSD are illustrated, including the genitalia of males and females, when available.

Taxonomic revision of the “ Pierella lamia species group” (Lepidoptera: Nymphalidae: Satyrinae) with descriptions of four new species from Brazil

Four new species of Pierella Westwood, 1851 from Brazil are described: P. angeloi Zacca, Siewert & Mielke sp. nov. from Maranhão, P. kesselringi Zacca, Siewert & Paluch sp. nov. from Paraíba, Pernambuco, Alagoas and Sergipe, P. nice Zacca, Siewert & Paluch sp. nov. from Bahia and P. keithbrowni Siewert, Zacca & Casagrande sp. nov. from Bahia, Espírito Santo, Rio de Janeiro, São Paulo, Paraná and Santa Catarina. Additionally, P. chalybaea Godman, 1905 stat. rest. and P. boliviana F.M. Brown, 1948 stat. nov. are recognized as valid species and not as subspecies of P. lamia (Sulzer, 1776), while P. l. colombiana Constantino & Salazar, 2007 syn. nov. is synonymized to the former. Lectotype and paralectotype of Papilio dyndimene Cramer, 1779 (a synonym of Pierella lamia) and Pierella chalybaea Godman, 1905 stat. rest. are designated. Habitus and illustrations of male and female genitalia are provided for all species, as well as a geographical distribution map.

Tosta Evans, 1953 (Lepidoptera: Hesperiidae): a new species from northeast Brazil, first description of the female of Tosta tosta Evans, 1953, and placement of the genus within Achlyodini.

A new species of Tosta Evans, 1953 from northeast Brazil is described: T. pseudospeculum Siewert, Mielke & Casagrande, sp. nov. Additionally, the female of Tosta tosta Evans, 1953, the type species of the genus, is described for the first time. Illustrations of adults and male and female genitalia for all studied species are provided, as well as a discussion of the placement of the genus within the tribe Achlyodini.