Stewart G. Molyneux

The stratigraphy, correlation, provenance and palaeogeography of the Skiddaw Group (Ordovician) in the English Lake District

A new lithostratigraphy is presented for the Skiddaw Group (lower Ordovician) of the English Lake District. Two stratigraphical belts are described. Five formations are defined in the Northern Fells Belt, ranging in age from Tremadoc to early Llanvirn. They are all mudstone or sandstone dominated, of turbidite origin; in ascending order they are named the Bitter Beck, Watch Hill, Hope Beck, Loweswater and Kirk Stile formations. Two formations are defined in the Central Fells Belt, ranging in age from late Arenig to Llanvirn. These are the Buttermere Formation - a major olistostrome deposit - overlain by the Tarn Moor Formation, consisting of turbidite mudstones with volcaniclastic turbidite sandstone beds. A revised graptolite and new acritarch biostratigraphy for the Skiddaw Group is presented with eight graptolite biozones and thirteen acritarch assemblages and sub-assemblages. The provenance of the group is assessed from detailed petrographical and geochemical work. This suggests derivation, in the early Ordovician, largely from an old inactive continental arc terrane lying to the south-east, with the appearance of juvenile volcanic material in the Llanvirn. Comparisons and correlations of the Skiddaw Group are made with the Isle of Man and eastern Ireland.

Ordovician organic-walled microphytoplankton (acritarch) distribution: the global scenario

Abstract A number of palaeobiogeographical models for Ordovician organic-walled microphytoplankton (acritarchs, prasinophytes, and related groups) have been published during the past 30 years. A modern synthesis of Ordovician acritarch palaeobiogeography, based on previously published acritarch ‘provinces’ and global distribution models, as well as new plots on recently compiled palaeogeographical maps is presented. Review of the literature and new plots indicate that a number of preliminary conclusions can be drawn. Following minor biogeographical differentiation of acritarch assemblages during the Cambrian, ‘provincialism’ started at the Cambrian–Ordovician boundary. In the late Tremadocian a warm-water assemblage, containing the genera Aryballomorpha , Athabascaella and Lua , but no diacrodians, seems to be limited to low-latitude localities such as Laurentia and North China. From the late Tremadocian and throughout most of the Arenig a peri-Gondwana acritarch assemblage with the easily recognisable taxa Arbusculidium filamentosum , Coryphidium , and Striatotheca is present on the southern margin of Gondwana, and its distribution corresponds almost exactly with that of the Calymenacean–Dalmanitacean trilobite fauna. It seems reasonable to consider the acritarchs of Baltica as belonging to a temperate-water ‘province’, which was probably not restricted to the palaeocontinent of Baltica but had a wider distribution at about the same latitude, as some of the elements recorded from Baltica also occur in South China and Argentina. The maximum separation of the continents during the Arenigian, reflected by a pronounced biogeographical differentiation of most Ordovician fossil groups, led to the development of geographically distinct acritarch assemblages. Data from the late Middle Ordovician and the Late Ordovician remain too poor to elucidate global palaeobiogeographical patterns. The biogeographical distribution of Ordovician acritarchs appears similar to that of the resting cysts of modern dinoflagellates, primarily controlled by latitude but also following the continental margins.

Late Ordovician Palynomorphs

ACRITARCHS Ordovician acritarchs have been recorded in five core samples collected between 2520 ft. and 3000 ft. in Well E1-81, and ten cutting samples taken between 12150 ft. and 13240 ft. in Well J1-81A. All the assemblages recovered are of Late Ordovician age; no Early Ordovician or Middle Ordovician assemblages have been identified. Investigations have so far concentrated on the acritarch assemblages from Well El-81. The highest three Ordovician samples from depths of 2520 to 2550 ft., 2552 to 2557 ft., and 2562 to 2567 ft., yielded similar assemblages which include Veryhachium irroratum, V. cf. lairdii, V. oklahomense?, V. subglobosum, V. trispinosum, Villosacapsula setosapellicula and a new species, Striatotheca sp. A. Navifusa similis? is represented by one specimen in the sample from 2552 to 2557 ft. Another specimen from the same sample is tentatively referred to Aremoricanium syringosagis. Specimens of Baltisphaeridium, Peteinosphaeridium, Leiofusa and Eupoikilofusa occur throughout the interval 2520 to 2567 ft. but are rare. Commonly occurring species include V. irroratum and V. setosapellicula. V. irroratum has been recorded from the Middle Ordovician of North America (Loeblich & Tappan, 1969) and the Caradoc of England (Turner, 1984) but Cramer & Diez (1979) maintain that it has its acme in the Ashgill. V. setosapellicula is common in the Sylvan Shale of Oklahoma (Loeblich, 1970) which is generally understood to be of Ashgill age, but is rare in the Eden Shale (Caradoc) of Indiana (Colbath, 1979) and in the type section of the Caradoc Series in Shropshire, England (Turner, 1984). . . .

THE ACRITARCH GENUS VERYHACHIUM DEUNFF 1954: TAXONOMIC EVALUATION AND FIRST APPEARANCE

Abstract Veryhachium Deunff 1954, originally described from the Ordovician of western France, is one of the most frequently recorded acritarch genera. Over 250 species and subspecies, from the Cambrian to the Neogene, have been attributed to the genus. This genus has a simple morphology; it displays a triangular, rectangular, or polygonal central vesicle, with a few, simple processes drawn out from the angles of the vesicle in a single plane, and sometimes with supplementary or auxillary processes arising from the vesicle body. Veryhachium has been emended and revised numerous times. The number of valid species is excessive: most are probably synonyms. To facilitate effective classification, only a few morphological categories should be retained. For the Lower Paleozoic, the use of two informal groups is proposed. These are the Veryhachium trispinosum group for triangular specimens, and the Veryhachium lairdii group for rectangular forms. Although generally abundant and widespread throughout the Phanerozoic, Veryhachium is of limited biostratigraphic, paleoecologic, or paleogeo-graphic value. However, its First Appearance Datum (FAD) is of great importance for Ordovician stratigraphy; the first Veryhachium morphotypes appear in the Tremadocian Stage, making the genus an important biostratigraphic marker.

The messaoudensis—trifidum acritarch assemblage and correlation of the base of Ordovician Stage 2 (Floian)

The Global Stratotype Section and Point (GSSP) for Stage 2 of the Ordovician System, now the Floian Stage and approximately equivalent to the lower and middle Arenig of England and Wales, is defined by the first appearance datum (FAD) of the graptolite Tetragraptus approximatus in the Diabasbrottet Quarry section at Mount Hunneberg, Sweden. One of the issues this raises is how to correlate the base of Stage 2 at the GSSP with areas and successions that do not contain a correlative graptolite fauna. The distinctive Cymatiogalea messaoudensis–Stelliferidium trifidum acritarch assemblage is present in the upper Tremadocian Araneograptus murrayi Graptolite Biozone of NW England and ranges across the Tremadocian–Stage 2 boundary there (the Tremadoc–Arenig boundary of Anglo-Welsh nomenclature). It also occurs widely at other high southern Ordovician palaeolatitudes around the margin of Gondwana, being reported from Ireland, Wales, the Isle of Man, Belgium, Germany, Spain and Turkey, and may also be present in Bohemia and Argentina (Eastern Cordillera). It therefore has the potential to contribute towards the recognition and correlation of the base of Stage 2 in those areas. Of particular interest are the First Appearance Datums of various taxa within the stratigraphical range of the messaoudensis–trifidum assemblage, notably that of Aureotesta clathrata simplex , which is considered to be close to the base of Stage 2 in NW England. Elements of the messaoudensis–trifidum assemblage also occur in Baltica, the palaeoplate on which the GSSP for the base of Stage 2 is located. However, many of the taxa used to subdivide the messaoudensis–trifidum assemblage around Gondwana have not been recorded from Baltica and may be restricted palaeobiogeographically to the Gondwanan margin. Furthermore, acritarch microfloras have not been reported from the Diabasbrottet Quarry section itself, and there are hiatuses across the base of Stage 2 in the two sections from Baltica considered in this paper. Hence, direct correlation of the base of Stage 2 between the GSSP and other sections using acritarchs is not yet possible. Nevertheless, some taxa, for example the genera Peteinosphaeridium and Rhopaliophora , are shown to have FADs at similar stratigraphical levels in the late Tremadocian Stage of both Baltica and Gondwana, and therefore have the potential to correlate time slices in the late Tremadocian Stage between palaeoplates.

The age and correlation of Skiddaw Group (early Ordovician) sediments in the Cross Fell inlier (northern England)

The Skiddaw Group in the Cross Fell inlier comprises the Catterpallot Formation of latest Tremadoc or earliest Arenig age, the Murton Formation of Arenig age, and the Kirkland Formation of early Llanvirn age. Each of these formations can be correlated with formations in the Skiddaw Group of the Lake District. The faulted contact of the Catterpallot and Kirkland formations is the probable extension of the Causey Pike Fault (CPF), which separates two distinct sequences in the Skiddaw inlier of the northern Lake District. Contrasts across the CPF in the Cross Fell inlier reflect those seen in the Skiddaw inlier. The CPF is a major basement structure, separating markedly different successions in the Ordovician strata of northern England.

Recycled acritarchs as provenance indicators; implications for Caledonian terrane reconstruction

Silurian successions deposited on the margins of the Iapetus Ocean in the English Lake District, Southern Uplands of Scotland and Ireland contain recycled Ordovician acritarchs. These include taxa known only from Avalonia and Gondwana, with other recycled acritarchs possibly derived from Baltica. There is no palynological evidence to indicate that Laurentiaconstituted a source area. The data presented herein suggest that provinciality in derived microfossils may be profitably exploited in provenance studies.

Preliminary account of Late Devonian palynomorph assemblages from the Bedford Shale and Berea Sandstone Formations of central Ohio, U.S.A.

Diverse assemblages of well preserved miospores and microplankton have been obtained from a series of samples collected across the boundary between the Bedford Shale and Berea Sandstone Formations in central Ohio. The miospores indicate a Late Devonian (pre-Carboniferous Tournaisian)* age for each of the samples. The microplankton support this age determination. A number of microplankton species not previously recorded from North America are present.

A revised correlation of the Cambrian rocks in the British Isles

This work reviews the correlation of the British and Irish Cambrian with the current (though incomplete) international standard for the Cambrian. Since the earlier edition of 1972, the basal and upper limits of the Cambrian system has been internationally agreed; so this account excludes Tremadocian rocks but includes some that were formerly considered Neoproerozoic. Half of the series and stage subdivisions are internationally agreed, but for the undefined divisions of the Cambrian the standard used here makes use of data from Avalonian successions. Since the first edition was published, almost every aspect of the Cambrian I the British Isles has been subjected to new study. Here, the plate tectonic make-up of the British Isles is reviewed, new radiometric ages and isotopic studies are summarized and the biostratigraphy is enhanced by the study of acritarchs, especially in the Irish successions

Chapter 23 Biogeography of early to mid Palaeozoic (Cambrian–Devonian) marine phytoplankton

Abstract Early to mid Palaeozoic marine phytoplankton are represented by acritarchs and associated forms, which had a global distribution from the early Cambrian to the early Carboniferous (Mississippian). Palaeozoic phytoplankton assemblages show varying degrees of cosmopolitanism and endemism through time. A high degree of cosmopolitanism was evidently characteristic of the Cambrian and much of the Late Ordovician, Silurian and Devonian, but provincialism was more marked in the Early Ordovician and Hirnantian (latest Ordovician), the latter at a time of major palaeoenvironmental perturbations. Distribution patterns of Palaeozoic phytoplankton are attributed to a number of interacting factors, including palaeolatitude, palaeotemperature, oceanic circulation patterns, the disposition of continents, differentiation between oceanic and cratonic (distal–proximal) assemblages, and sedimentary environments and facies. There are indications that biogeographical ranges of taxa shift over time. Moving our understanding of Palaeozoic phytoplankton biogeography forward requires (1) targeted investigation of regions and time periods for which no or little data exist, (2) quantitative analysis of data to investigate how similarity between regions varies through time and how this might correlate with other datasets such as carbon isotope stratigraphy or sea-level, and (3) rigorous application of well-defined time slices to compare coeval assemblages, at least within the limits of resolution.